Alfred G. Knudson

Alfred George Knudson, Jr. M.D., Ph.D. (August 9, 1922 – July 10, 2016) was an American geneticist. He worked in cancer genetics. He was known for the formulation of the Knudson hypothesis in 1971. It explains the effects of mutation on carcinogenesis (the development of cancer).
Knudson died on July 10, 2016 at his home in Philadelphia, Pennsylvania from a long-illness at the age of 93.