text:
List of organisms by chromosome count
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Karyotype of a human being, showing 22 pair of autosomal chromosomes and both the XX female and the XY male possibilities for the pair of allosomic (sex) chromosome
Fusion of ancestral chromosomes left distinctive remnants of telomeres, and a vestigial centromere. As other non-human extant hominidae have 48 chromosomes it is believed that the human chromosome 2 is the end result of the merging of two chromosomes.[1]
The list of organisms by chromosome count describes ploidy or numbers of chromosomes in the cells of various plants, animals, protists, and other living organisms. This number, along with the visual appearance of the chromosome, is known as the karyotype,[2][3][4] and can be found by looking at the chromosomes through a microscope. Attention is paid to their length, the position of the centromeres, banding pattern, any differences between the sex chromosomes, and any other physical characteristics.[5] The preparation and study of karyotypes is part of cytogenetics.
Animals
Plants
Other Eukaryotes
Organism
(Scientific name)
Chromosome number
Picture
Karyotype
Notes
Source
Jack jumper ant
(Myrmecia pilosula)
02/1 !2/1 2 for females, males are haploid and thus have 1; smallest number possible. Other ant species have more chromosomes.[6] [6]
Australian daisy
(Brachyscome dichromosomatica)
12 !12 This species can have more B chromosomes than A chromosomes at times, but 2n=4. [7]
Spider mite (Tetranychidae)
04 !4–14 Spider mites (family Tetranychidae) are typically haplodiploidy (males are haploid, while females are diploid)[8] [8]
Yellowfever mosquito
(Aedes aegypti)
06 !6 The 2n=6 chromosome number is conserved in the entire family Culicidae, except in Chagasia bathana, which has 2n=8.[9] [9]
Indian muntjac
(Muntiacus muntjak)
06/7 !6/7 2n = 6 for females and 7 for males. The lowest diploid chromosomal number in mammals.[10] [11]
Hieracium
08 !8
Fruit fly
(Drosophila melanogaster)
08 !8 6 autosomal and 2 allosomic (sex) [12]
Macrostomum lignano
(Macrostomum lignano)
08 !8 [13]
Thale cress
(Arabidopsis thaliana)
10
Swamp wallaby
(Wallabia bicolor)
10/11 11 for male, 10 for female [14]
Nematode
(Caenorhabditis elegans)
12/11 12 for hermaphrodites, 11 for males
Spinach
(Spinacia oleracea)
12 [15]
Broad bean
(Vicia faba)
12 [16]
Yellow dung fly
(Scathophaga stercoraria)
12 10 autosomal and 2 allosomic (sex) chromosomes. Males have XY sex chromosomes and females have XX sex chromosomes. The sex chromosomes are the largest chromosomes and constitute 30% of the total length of the diploid set in females and about 25% in males.[17] [17]
Slime mold
(Dictyostelium discoideum)
12 [18]
Cucumber
(Cucumis sativus)
14 [19]
Tasmanian devil
(Sarcophilus harrisii)
14
Rye
(Secale cereale)
14 [20]
Pea
(Pisum sativum)
14 [20]
Barley
(Hordeum vulgare)
14 [21]
Aloe vera
14 The diploid chromosome number is 2n = 14 with four pair of long acrocentric chromosomes ranging from 14.4 μm to 17.9 μm and three pair of short sub metacentric chromosomes ranging from 4.6 μm to 5.4 μm.[22] [22]
Koala
(Phascolarctos cinereus)
16
Kangaroo
16 This includes several members of genus Macropus, but not the red kangaroo (M. rufus, 20) [23]
Schistosoma mansoni
16 2n=16. 7 autosomal pairs and ZW sex-determination pair.[24] [24]
Welsh onion
(Allium Fistulosum)
16 [25]
Garlic
(Allium sativum)
16 [25]
Itch mite
(Sarcoptes scabiei)
17 !17/18 According to the observation of embryonic cells of egg, chromosome number of the itch mite is either 17 or 18. While the cause for the disparate numbers is unknown, it may arise because of an XO sex determination mechanism, where males (2n=17) lack the sex chromosome and therefore have one less chromosome than the female (2n=18).[26] [26]
Radish
(Raphanus sativus)
18 [20]
Carrot
(Daucus carota)
18 The genus Daucus includes around 25 species. D. carota has nine chromosome pairs (2n = 2x = 18). D. capillifolius, D. sahariensis and D. syrticus are the other members of the genus with 2n = 18, whereas D. muricatus (2n = 20) and D. pusillus (2n = 22) have a slightly higher chromosome number. A few polyploid species as for example D. glochidiatus (2n = 4x = 44) and D. montanus (2n = 6x = 66) also exist.[27] [27]
Cabbage
(Brassica oleracea)
18 Broccoli, cabbage, kale, kohlrabi, brussels sprouts, and cauliflower are all the same species and have the same chromosome number.[20] [20]
Citrus
(Citrus x)
18 Chromosome number of the genus Citrus, which including lemons, oranges, grapefruit, pomelo and limes, is 2n = 18.[28] [29]
Passion fruit
(Passiflora edulis)
18 [30]
Setaria viridis
(Setaria viridis)
18 [31]
Maize
(Zea mays)
20 [20]
Cannabis
(Cannabis sativa)
20
Western clawed frog
(Xenopus tropicalis)
20 [32]
Australian pitcher plant
(Cephalotus follicularis)
20 [33]
Cacao
(Theobroma cacao)
20 [34]
Eucalyptus
(Eucalyptus)
22 Although some contradictory cases have been reported, the large homogeneity of the chromosome number 2n = 22 is now known for 135 (33.5%) distinct species among genus Eucalyptus.[35] [36]
Virginia opossum
(Didelphis virginiana)
22 [37]
Bean
(Phaseolus sp.)
22 All species in the genus Phaseolus have the same chromosome number, including common bean (P. vulgaris), runner bean (P. coccineus), tepary bean (P. acutifolius) and lima bean (P. lunatus).[20] [20]
Snail
24
Melon
(Cucumis melo)
24 [38]
Rice
(Oryza sativa)
24 [20]
Silverleaf nightshade
(Solanum elaeagnifolium)
24 [39]
Sweet chestnut
(Castanea sativa)
24 [40]
Tomato
(Solanum lycopersicum)
24 [41]
European beech
(Fagus sylvatica)
24 [42]
Bittersweet nightshade
(Solanum dulcamara)
24 [43][44]
Cork oak
(Quercus suber)
24 [45]
Edible frog
(Pelophylax kl. esculentus)
26 Edible frog is the fertile hybrid of the pool frog and the marsh frog.[46] [47]
Axolotl
(Ambystoma mexicanum)
28 [48]
Bed bug
(Cimex lectularius)
29–47 26 autosomes and varying number of the sex chromosomes from three (X1X2Y) to 21 (X1X2Y+18 extra Xs).[49] [49]
Pill millipede
(Arthrosphaera magna attems)
30 [50]
Giraffe
(Giraffa camelopardalis)
30 [51]
American mink
(Neovison vison)
30
Pistachio
(Pistacia vera)
30 [52]
Yeast
(Saccharomyces cerivisiae)
32
European honey bee
(Apis mellifera)
32/16 32 for females (2n = 32), males are haploid and thus have 16 (1n =16).[53] [53]
American badger
(Taxidea taxus)
32
Alfalfa
(Medicago sativa)
32 Cultivated alfalfa is tetraploid, with 2n=4x=32. Wild relatives have 2n=16.[20]:165 [20]
Red fox
(Vulpes vulpes)
34 Plus 3-5 microsomes. [54]
Sunflower
(Helianthus annuus)
34 [55]
Porcupine
(Erethizon dorsatum)
34 [56]
Globe artichoke
(Cynara cardunculus var. scolymus)
34 [57]
Yellow mongoose
(Cynictis penicillata)
36
Tibetan sand fox
(Vulpes ferrilata)
36
Starfish
(Asteroidea)
36
Red panda
(Ailurus fulgens)
36
Meerkat
(Suricata suricatta)
36
Cassava
(Manihot esculenta)
36 [58]
Long-nosed cusimanse
(crossarchus obscurus)
36
Earthworm
(Lumbricus terrestris)
36
African clawed frog
(Xenopus laevis)
36 [32]
Waterwheel plant
(Aldrovanda vesiculosa)
38 [33]
Tiger
(Panthera tigris)
38
Sea otter
(Enhydra lutris)
38
Sable
(Martes zibellina)
38
Raccoon
(Procyon lotor)
38 [59]
Pine marten
(Martes martes)
38
Pig
(Sus)
38
Oriental small-clawed otter
(Aonyx cinerea)
38
Lion
(Panthera leo)
38
Fisher
38 a type of marten
European mink
(Mustela lutreola)
38
Coatimundi
38
Cat
(Felis silvestris catus)
38
Beech marten
(Martes foina)
38
Baja California ratsnake
(Bogertophis rosaliae)
38 [60]
American marten
(Martes americana)
38
Trans-Pecos ratsnake
(Bogertophis subocularis)
40 [61]
Mouse
(Mus musculus)
40 [62]
Mango
(Mangifera indica)
40 [20]
Hyena
(Hyaenidae)
40
Ferret
(Mustela putorius furo)
40
European polecat
(Mustela putorius)
40
Beaver (American)
(Castor canadensis)
40
Peanut
(Arachis hypogaea)
40 Cultivated peanut is an allotetraploid (2n = 4x = 40). Its closest relatives are the diploid (2n = 2x = 20).[63] [63]
Wolverine
(Gulo gulo)
42
Wheat
(Triticum aestivum)
42 This is a hexaploid with 2n=6x=42. Durum wheat is Triticum turgidum var. durum, and is a tetraploid with 2n=4x=28.[20] [20]
Rhesus monkey
(Macaca mulatta)
42 [64]
Rat
(Rattus norvegicus)
42 [65]
Oats
(Avena sativa)
42 This is a hexaploid with 2n=6x=42. Diploid and tetraploid cultivated species also exist.[20] [20]
Giant panda
(Ailuropoda melanoleuca)
42
Fossa
(Cryptoprocta ferox)
42
European rabbit
(Oryctolagus cuniculus)
44
Eurasian badger
(Meles meles)
44
Moon jellyfish
(Aurelia aurita)
44 [66]
Dolphin
(Delphinidae Delphi)
44
Coffea arabica
(Coffea arabica)
44 Out of the 103 species in the genus Coffea, arabica coffee is the only tetraploid species (2n = 4x = 44), the remaining species being diploid with 2n = 2x = 22.[67]
Sable antelope
(Hippotragus niger)
46
Reeves's muntjac
(Muntiacus reevesi)
46
Human
(Homo sapiens)
46 44 autosomal. and 2 allosomic (sex) [68]
Parhyale hawaiensis
46 [69]
Water buffalo (river type)
(Bubalus bubalis)
48
Tobacco
(Nicotiana tabacum)
48 Cultivated species N. tabacum is an amphidiploid (2n=4x=48) evolved through the interspecific hybridization of the ancestors of N. sylvestris (2n=2x=24, maternal donor) and N. tomentosiformis (2n=2x=24, paternal donor) about 200,000 years ago.[70] [70]
Potato
(Solanum tuberosum)
48 This is for common potato Solanum tuberosum (tetraploid, 2n = 4x = 48). Other cultivated potato species may be diploid (2n = 2x = 24), triploid (2n = 3x = 36), tetraploid (2n = 4x = 48), or pentaploid (2n = 5x = 60).[71] Wild relatives mostly have 2n=24.[20] [71]
Orangutan
(Pongo)
48
Hare
(Lepus)
48 [72][73]
Gorilla
(Gorilla)
48
Deer mouse
(Peromyscus maniculatus)
48
Chimpanzee
(Pan troglodytes)
48 [74]
Beaver (Eurasian)
(Castor fiber)
48
Zebrafish
(Danio rerio)
50 [75]
Water buffalo (swamp type)
(Bubalus bubalis)
50
Striped skunk
(Mephitis mephitis)
50
Pineapple
(Ananas comosus)
50 [20]
Kit fox
(Vulpes macrotis)
50
Spectacled bear
(Tremarctos ornatus)
52
Platypus
(Ornithorhynchus anatinus)
52 Ten sex chromosomes. Males have X1Y1X2Y2X3Y3X4Y4X5Y5, females have X1X1X2X2X3X3X4X4X5X5.[76] [77]
Cotton
(Gossypium hirsutum)
52 This is for the cultivated species G. hirsutum (allotetraploid, 2n=4x=52). This species accounts for 90% of the world cotton production. Among 50 species in the genus Gossypium, 45 are diploid (2n = 2x = 26) and 5 are allotetraploid (2n = 4x = 52).[78] [78]
Sheep
(Ovis orientalis aries)
54
Hyrax
(Hyracoidea)
54 !54 Hyraxes were considered to be the closest living relatives of Elephants,[79] but sirenians have been found to be more closely related to elephants. [80]
Raccoon dog
(Nyctereutes procyonoides procyonoides)
54 This number is for Chinese raccoon dog (N. p. procyonoides), 2n=54+B(0–4). On the other hand, Japanese raccoon dog (N. p. viverrinus) with 2n=38+B(0–8). Here, B represents B chromosome and its variation in the number between individuals.[81][82] [81]
Capuchin monkey
(Cebus x)
54 [83]
Silkworm
(Bombyx mori)
56 This is for the species mulberry silkworm, B. mori (2n=56). Probably more than 99% of the world's commercial silk today come from this species.[84] Other silk producing moths, called non-mulberry silkworms, have various chromosome numbers. (e.g. Samia cynthia with 2n=25–28,[85] Antheraea pernyi with 2n=98.[86]) [87]
Strawberry
(Fragaria)
56 This number is octoploid, main cultivated species Fragaria × ananassa (2n = 8x = 56). In genus Fragaria, basic chromosome number is seven (x = 7) and multiple levels of ploidy, ranging from diploid (2n = 2x = 14) to decaploid (F. iturupensis, 2n = 10x = 70), are known.[88] [88]
Gaur
(Bos gaurus)
56
Elephant
(Elephantidae)
56
Woolly mammoth
(Mammuthus primigenius)
58 extinct; tissue from a frozen carcass
Yak
(Bos mutus)
60
Goat
(Capra aegagrus hircus)
60
Cow/Bull
(Bos primigenius)
60
American bison
(Bison bison)
60
Bengal fox
(Vulpes bengalensis)
60
Gypsy moth
(Lymantria dispar dispar)
62
Donkey
(Equus africanus asinus)
62
Scarlet macaw
(Ara macao)
62–64 [89]
Mule
63 semi-infertile (odd number of chromosomes – between donkey (62) and horse (64) makes meiosis much more difficult)
Guinea pig
(Cavia porcellus)
64
Spotted skunk
(Spilogale x)
64
Horse
(Equus ferus caballus)
64
Fennec fox
(Vulpes zerda)
64 [54]
Echidna
63/64 63 (X1Y1X2Y2X3Y3X4Y4X5, male) and 64 (X1X1X2X2X3X3X4X4X5X5, female)[90]
Chinchilla
(Chinchilla lanigera)
64 [56]
Nine-banded armadillo
(Dasypus novemcinctus)
64 [91]
Gray fox
(Urocyon cinereoargenteus)
66 [54]
Red deer
(Cervus elaphus)
68
Elk (Wapiti)
(Cervus canadensis)
68
Roadside hawk
(Rupornis magnirostris)
68 [92]
White-tailed deer
(Odocoileus virginianus)
70
Black nightshade
(Solanum nigrum)
72 [93]
Bat-eared fox
(Otocyon megalotis)
72 [54]
Sun bear
(Helarctos malayanus)
74
Sloth bear
(Melursus ursinus)
74
Polar bear
(Ursus maritimus)
74
Brown bear
(Ursus arctos)
74
Asiatic black bear
(Ursus thibetanus)
74
American black bear
(Ursus americanus)
74
Maned wolf
(Chrysocyon brachyurus)
76
Gray wolf
(Canis lupus)
78
Golden jackal
(Canis aureus)
78 [54]
Dove
(Columbidae)
78 Based on African collared dove [94]
Dog
(Canis lupus familiaris)
78 Normal dog karyotype is composed of 38 pairs of acrocentric autosomes and two metacentric sex chromosomes.[95][96] [97]
Dingo
(Canis lupus dingo)
78 [54]
Dhole
(Cuon alpinus)
78
Coyote
(Canis latrans)
78 [54]
Chicken
(Gallus gallus domesticus)
78
African wild dog
(Lycaon pictus)
78 [54]
Tropical pitcher plant
(Nepenthes rafflesiana)
78 [33]
Turkey
(Meles)
80 [98]
Sugarcane
(Saccharum officinarum)
80 This is for S. officinarum (octoploid, 2n = 8× = 80).[99] About 70% of the world’s sugar comes from this species.[100] Other species in the genus Saccharum, collectively known as sugarcane, have chromosome numbers in the range 2n=40–128.[101] [99]
Pigeon
(Columbidae)
80 [102]
Great white shark
(Carcharodon carcharias)
82 [103]
Hedgehog genus Erinaceus (woodland hedgehogs)
88
Moonworts
(Botrychium)
90
Hedgehog Genus Atelerix (African hedgehogs)
90
Grape fern
(Sceptridium)
90
Pittier's crab-eating rat
(Ichthyomys pittieri)
92 Previously thought to be the highest number in mammals, tied with Anotomys leander. [104]
Shrimp
(Penaeus semisulcatus)
86–92 [105]
Aquatic rat
(Anotomys leander)
92 Previously thought to be the highest number in mammals, tied with Ichthyomys pittieri. [104]
Kamraj (fern)
(Helminthostachys zeylanica)
94
Carp
(Carassius carassius)
:100 !100 [106]
Red viscacha rat
(Tympanoctomys barrerae)
:102 !102 Highest number known in mammals, thought to be a tetraploid[107] or allotetraploid.[108] [109]
Walking catfish
(Clarias batrachus)
104 [110]
American paddlefish
(Polyodon spathula)
120 [111]
Northern lamprey
(Petromyzontinae)
:164 !174 [112]
Rattlesnake fern
(Botrypus virginianus)
:184 !184 [113]
Red king crab
(Paralithodes camtschaticus)
208
Field horsetail
(Equisetum arvense)
:216 !216
Adders-tongue
(Ophioglossum)
:240 !240 n=120–720 with a high degree of polyploidization[114] n=720 in hexaploid species O. reticulatum. A google book search for "ophioglossum reticulatum chromosomes" returns values of 2n=768 and 2n=1260, though these sources may be wp:circular and unreliable.
Agrodiaetus butterfly
(Agrodiaetus shahrami)
:268 !268 This insect has one of the highest chromosome numbers among all animals. [115]
Black mulberry
(Morus nigra)
:308 !308 In this case, the chromosome number is docosaploid, or 22 times the basic number. [116]
Atlas blue
(Polyommatus atlantica)
:448 !448-452 2n = circa 448–452. Highest number of chromosomes in the non-polyploid eukaryotic organisms.[117] [117]
Ciliated protozoa
(Tetrahymena thermophila)
10 (in micronucleus) 50x = 12,500 (in macronucleus, except minichromosomes)
10,000x = 10,000 (macronuclear minichromosomes)[118]
Ciliated protozoa
(Oxytricha trifallax)
15,600 MAC chromosomes × 1900 ploidy level =
29.64 × 106 chromosomes Macronuclear "nanochromosomes"; ampliploid. [119][120][121]
See also[edit]
Ecology portal
References[edit]
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^ Estienne C. Swart; John R. Bracht; Vincent Magrini; Patrick Minx; Xiao Chen; Yi Zhou; Jaspreet S. Khurana; Aaron D. Goldman; Mariusz Nowacki; Klaas Schotanus; Seolkyoung Jung; Robert S. Fulton; Amy Ly; Sean McGrath; Kevin Haub; Jessica L. Wiggins; Donna Storton; John C. Matese; Lance Parsons; Wei-Jen Chang; Michael S. Bowen; Nicholas A. Stover; Thomas A. Jones; Sean R. Eddy; Glenn A. Herrick; Thomas G. Doak; Richard K. Wilson; Elaine R. Mardis; Laura F. Landweber (2013-01-29). "The Oxytricha trifallax Macronuclear Genome: A Complex Eukaryotic Genome with 16,000 Tiny Chromosomes". PLOS Biology. 11 (1): e1001473. doi:10.1371/journal.pbio.1001473. PMC 3558436  . PMID 23382650. Retrieved 2014-04-29.
^ "You Have 46 Chromosomes. This Pond Creature Has 15,600", National Geographic, [3].
External links[edit]
List of pages in English from Russian bionet site
The dog through evolution
Shared synteny of human chromosome 17 loci in Canids
An atlas of the chromosome numbers in animals (1951); PDF downloads of each chapter
Bell, G. (1982). The Masterpiece of Nature: The Evolution and Genetics of Sexuality (University of California Press, Berkeley), p. 450, [4] (table with a compilation of haploid chromosome number of many algae and protozoa, in column "HAP").
Nuismer, S.; Otto, S.P. (2004). "Host-parasite interactions and the evolution of ploidy". Proc. Natl. Acad. Sci. USA. 101 (30): 11036–11039. doi:10.1073/pnas.0403151101. (Supporting Data Set, with information on ploidy level and number of chromosomes of several protists)
v
t
e
Cytogenetics: chromosomes
Basic
concepts
Karyotype
Ploidy
Genetic material/Genome
Chromatin
Euchromatin
Heterochromatin
Chromosome
Chromatid
Nucleosome
Nuclear organization
Types
Autosome/Sex chromosome (or allosome or heterosome)
Macrochromosome/Microchromosome
Circular chromosome/Linear chromosome
Extra chromosome (or accessory chromosome)
Supernumerary chromosome
A chromosome/B chromosome
Lampbrush chromosome
Polytene chromosome
Dinoflagellate chromosomes
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Satellite chromosome
Centromere position
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Centromere number
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Processes
and evolution
Mitosis
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Chromosomal inversion
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Aneuploidy
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Structures
Telomere: Telomere-binding protein (TINF2)
Protamine
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H3
H4
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C1
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F
H
I
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K
M
N
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Q
T
See also
Extrachromosomal DNA
Plasmid
List of organisms by chromosome count
List of chromosome lengths for various organisms
List of sequenced genomes
International System for Human Cytogenetic Nomenclature
Retrieved from "https://en.wikipedia.org/w/index.php?title=List_of_organisms_by_chromosome_count&oldid=834467484"
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Performance work statement
From Wikipedia, the free encyclopedia
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Performance work statement (or PWS) is to summarize the work that needs to be done for a contract (e.g., with the U.S. Department of Defense).
Include at least the following sections for an acquisition: scope; applicable documents; performance requirements/tasks; and, contractor quality assurance.
Provide the following information: customer contact information; period of performance; place of performance; background; scope; applicable directives; government furnished property/equipment/information; and, performance requirements.
References[edit]
https://www.atsc.army.mil/TADLP/content/pws/index.asp
https://dap.dau.mil/acquipedia/Pages/ArticleDetails.aspx?aid=488854b0-d8ee-4e32-aa3e-301d2ac8ffca
Retrieved from "https://en.wikipedia.org/w/index.php?title=Performance_work_statement&oldid=842448422"
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Attack on Titan (video game)
From Wikipedia, the free encyclopedia
Jump to: navigation, search
Attack on Titan
Developer(s)
Omega Force
Publisher(s)
Koei Tecmo
Director(s)
Tomoyuki Kitamura
Producer(s)
Hisashi Koinuma
Designer(s)
Kotaro Hirata
Programmer(s)
Eita Karasawa
Writer(s)
Ryohei Hayashi
Composer(s)
Shinichiro Nakamura
Yugen Umemura
Junya Ishiguro
Series
Attack on Titan
Platform(s)
Microsoft Windows
PlayStation 3
PlayStation 4
PlayStation Vita
Xbox One
Release
JP: February 18, 2016
PAL: August 26, 2016[1]
NA: August 30, 2016[2]
Genre(s)
Action, hack and slash
Mode(s)
Single-player, multiplayer
Attack on Titan[2] (Japanese: 進撃の巨人, Hepburn: Shingeki no Kyojin), known in Europe as A.O.T.: Wings of Freedom,[1] is an action hack and slash video game based on Hajime Isayama's manga series of the same name released for PlayStation 3, PlayStation 4, and PlayStation Vita in Japan. It was released on February 18, 2016, in Japan.[3] On April 7, Koei Tecmo announced a Western release in North America and Europe for August 2016.[1][2] Microsoft Windows and Xbox One versions were also revealed.[2] On August 21, 2017, Koei Tecmo America confirmed that Attack on Titan 2 has been set for a 2018 release.[4]
Contents
[hide]
1 Plot
2 Gameplay
3 Characters
4 Development
5 Reception
6 Sequel
7 References
Plot[edit]
The game retells key moments from the Attack on Titan manga (1 through 19-), in addition to new scenarios involving the key characters.[5]
In 845, when Eren was just 10 years old, the "Colossal Titan" suddenly appeared. Large enough to breach the Wall, it allowed other Titans to rampage through the city. One of them ate Carla, Eren's mother. With the so-called "Armored Titan" now preventing access, Wall Maria was abandoned and humanity were forced to fall back to Wall Rose. Having lost his family, his home and his dreams, Eren swore revenge on the Titans. Aiming to become a member of the Scout Regiment, he joined the "104th Cadet Corps."
Three years after Eren enlisted, the "Colossal Titan" suddenly appeared again. Eren tried to fight it alone but failed to defeat it. He returned to headquarters and then takes a squad out himself, but the mission was a failure with Eren suffering a terrible injury to his leg and many soldiers being killed. When Armin was about to be eaten by a Titan, Eren sacrifices his life to save him. Immediately before the Trost District Wall was destroyed by the Colossal Titan, elite members of the Scout Regiment were on a mission outside the Wall. The principle role of the Scout Regiment was to scout beyond the Wall, seeking a way to take back Wall Maria which was lost 5 years previously. Setting up supply bases in destroyed towns and villages, they sought to secure a route for a large force to make their way to the Shiganshina District, where the Wall was destroyed. Fighting with the Titans outside the wall as they go, they sought to secure the route while making new discoveries along the way.
The brave resistance offered by the Garrison Regiment, combined with the return of the Scout Regiment, was enough to keep Wall Rose safe from the Titans. Within the Walls, however, the same chaos descended as was experienced when Wall Maria was lost 5 years previously. Not only has the Colossal Titan attacked humanity again, but the damaged Wall has been repaired - unbelievably - by the "power of the Titans." Is Eren an ally for humanity or just another enemy? As the debate continues within the Walls, the Scout Regiment - newly joined by Eren - begin their 57th mission to the outside world.[6]
Gameplay[edit]
The game's visuals matches that of the anime series and it features one player fighting multiple enemies. The 3D Maneuvering Gear can be used to ascend to high areas.[7]
Characters[edit]
See also: List of Attack on Titan characters
The game features ten playable characters. Initially, only three can be selected. But as players progress through Attack (story) mode, more characters are unlocked.
Eren Jaeger
Mikasa Ackerman
Armin Arlert
Jean Kirschtein
Connie Springer
Sasha Braus
Krista Lenz
Levi
Zoë Hanji
Erwin Smith
Development[edit]
The game was published by Koei Tecmo and developed by Omega Force, and was first announced at Gamescom.[8] A new trailer was unveiled on November 6, 2015, in addition to details about the Collector's Edition.[9] It was released for the PlayStation 3, PlayStation 4, and PlayStation Vita in Japan.[10] Subsequent versions for Xbox One and Microsoft Windows were announced following its Western announcement by Koei Tecmo.[2]
Reception[edit]
Reception
Review scores
Publication
Score
GameSpot 6/10[11]
IGN 7.3/10[12]
Attack on Titan sold approximately 150,682 copies in its first week in Japan,[13] outselling Street Fighter V during that week.[14] As of April 28, Koei Tecmo announced it had sold 280,000 copies in total in Japan.[15] As of Koei Tecmo's 2016/17 FY report, the game has sold in 350,000 copies in the West and a cumulative 700,000 copies worldwide. [16]
Attack on Titan received polarizing positive reviews. Famitsu's four reviewers awarded it consecutive scores of 8/9/9/8, which added up to an overall score of 34/40.[17] PlayStation LifeStyle awarded it a score of 8.0 out of 10, saying "Attack on Titan is an excellent example of a passion project made right."[18] However, in other places, it got mixed to negative reviews.
Sequel[edit]
A sequel game to Koei Tecmo's Attack on Titan (titled Attack on Titan 2) was announced in August 2017 and was released in March 2018.[19]
References[edit]
^ Jump up to: a b c "A.O.T. WINGS OF FREEDOM - ACTION TRAILER". YouTube. KOEI TECMO EUROPE LTD. Retrieved April 7, 2016.
^ Jump up to: a b c d e "Attack on Titan - Announcement Trailer". YouTube. KOEI TECMO AMERICA. Retrieved April 7, 2016.
Jump up ^ Miller, KK (December 1, 2015). "New 'Attack on Titan' game release date announced". Japan Today. Retrieved December 3, 2015.
Jump up ^ Romano, Sal (August 21, 2017). "Koei Tecmo and Omega Force announce Attack on Titan 2". Gematsu. Retrieved August 21, 2017.
Jump up ^ Sato (August 21, 2015). "Koei Tecmo's Attack On Titan Game Includes The Entire Story From The Anime". Siliconera. Retrieved December 3, 2015.
Jump up ^ "Attack on Titan". Attack on Titan Official Site. Retrieved 2017-03-15.
Jump up ^ Barder, Ollie (August 20, 2015). "Upcoming 'Attack On Titan' Game Looks A Lot Like The Anime". Forbes. Retrieved December 3, 2015.
Jump up ^ Gelmini, David (August 6, 2015). "Koei to Publish New Attack on Titan Game". Dread Central. Retrieved December 3, 2015.
Jump up ^ Barder, Ollie (6 November 2015). "'Attack On Titan' Game Gets New Trailer And Treasure Box Collector's Edition". Forbes. Retrieved December 3, 2015.
Jump up ^ Sato (November 26, 2015). "Attack on Titan To Release In Japan On February 2016 For PS4, PS3, And Vita". Siliconera. Retrieved December 3, 2015.
Jump up ^ Corriea, Ray (August 26, 2016). "Attack on Titan Review". GameSpot. Retrieved February 25, 2017.
Jump up ^ Sanchez, Miranda (August 26, 2016). "Attack on Titan Review". IGN. Retrieved February 25, 2017.
Jump up ^ http://www.siliconera.com/2016/02/24/week-sales-kancolle-kai-attack-titan-street-fighter-v-take-charts/
Jump up ^ http://www.playstationlifestyle.net/2016/02/24/japanese-sales-chart-attack-titan-outsells-street-fighter-v-ps4-remains-1st/
Jump up ^ http://www.dualshockers.com/2016/04/28/total-sales-for-dead-or-alive-xtreme-3-attack-on-titan-and-more-announced-by-koei-tecmo/
Jump up ^ https://zhugeex.com/2016/11/koei-tecmo-h1-fy201617-report-attack-on-titan-sell-in-exceeds-700k-worldwide/
Jump up ^ Romano, Sal (February 9, 2016). "Famitsu Review Scores: Issue 1419 (PS4)". Gematsu. Retrieved February 25, 2017.
Jump up ^ Wood, Chandler (August 26, 2016). "Attack on Titan Review – Licensed to Kill (PS4)". Retrieved February 25, 2017.
Jump up ^ Romano, Sal (October 24, 2017). "Koei Tecmo's Attack on Titan 2 launches March 2018 in North America and Europe". Gematsu. Retrieved February 10, 2018.
Video games portal
2010s portal
[hide]
v
t
e
Attack on Titan by Hajime Isayama
Media
Chapters
Episodes
Before the Fall
Junior High
Lost Girls
No Regrets
Spoof on Titan
Harsh Mistress of the City
Live-action
Attack on Titan
Counter Rockets
Video games
Humanity in Chains
Attack on Titan
Attack on Titan 2
Related
Characters
Eren Yeager
"Jiyū e no Shingeki" by Linked Horizon
Retrieved from "https://en.wikipedia.org/w/index.php?title=Attack_on_Titan_(video_game)&oldid=834337997"
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