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Cat
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This article is about the cat species that is commonly kept as a pet. For the cat family, see Felidae. For other uses, see Cat (disambiguation) and Cats (disambiguation).
For technical reasons, "Cat #1" redirects here. For that album, see Cat 1 (album).
Domestic cat[1]
Various types of domestic cat
Conservation status
Domesticated
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Carnivora
Suborder: Feliformia
Family: Felidae
Subfamily: Felinae
Genus: Felis
Species: F. silvestris
Subspecies: F. s. catus
Trinomial name
Felis silvestris catus
Linnaeus, 1758[2]
Synonyms
Felis catus (original combination)[3]
Felis catus domestica (invalid junior synonym)[4]
The domestic cat (Felis silvestris catus or Felis catus)[1][5] is a small, typically furry, carnivorous mammal. They are often called house cats[6] when kept as indoor pets or simply cats when there is no need to distinguish them from other felids and felines. They are often valued by humans for companionship and for their ability to hunt vermin. There are more than seventy cat breeds recognized by various cat registries.
Cats are similar in anatomy to the other felids, with a strong flexible body, quick reflexes, sharp retractable claws and teeth adapted to killing small prey. Cat senses fit a crepuscular and predatory ecological niche. Cats can hear sounds too faint or too high in frequency for human ears, such as those made by mice and other small animals. They can see in near darkness. Like most other mammals, cats have poorer color vision and a better sense of smell than humans. Cats, despite being solitary hunters, are a social species, and cat communication includes the use of a variety of vocalizations (mewing, purring, trilling, hissing, growling and grunting) as well as cat pheromones and types of cat-specific body language.[7]
Cats have a high breeding rate.[8] Under controlled breeding, they can be bred and shown as registered pedigree pets, a hobby known as cat fancy. Failure to control the breeding of pet cats by neutering, as well as the abandonment of former household pets, has resulted in large numbers of feral cats worldwide, requiring population control.[9] In certain areas outside cats' native range, this has contributed, along with habitat destruction and other factors, to the extinction of many bird species. Cats have been known to extirpate a bird species within specific regions and may have contributed to the extinction of isolated island populations.[10] Cats are thought to be primarily responsible for the extinction of 33 species of birds since the 1600s,[better source needed] and the presence of feral and free-ranging cats makes some otherwise suitable locations unsuitable for attempted species reintroduction.[11]
Because cats were venerated in ancient Egypt, they were commonly believed to have been domesticated there,[12] but there may have been instances of domestication as early as the Neolithic from around 9,500 years ago (7500 BC).[13] A genetic study in 2007[14] concluded that all domestic cats are descended from Near Eastern wildcats, having diverged around 8000 BC in the Middle East.[12][15] A 2016 study found that leopard cats were undergoing domestication independently in China around 5500 BC, though this line of partially domesticated cats leaves no trace in the domesticated populations of today.[16][17] A 2017 study confirmed that domestic cats are descendants of those first domesticated by farmers in the Near East around 9,000 years ago.[18][19]
As of a 2007 study, cats are the second-most popular pet in the U.S. by number of pets owned, behind freshwater fish.[20] In a 2010 study, they were ranked the third-most popular pet in the UK, after fish and dogs, with around 8 million being owned.[21]
Contents
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1 Taxonomy and evolution
2 Nomenclature and etymology
3 Biology
3.1 Anatomy
3.2 Physiology
3.2.1 Nutrition
3.3 Senses
3.4 Health
3.4.1 Diseases
3.5 Genetics
4 Behavior
4.1 Sociability
4.2 Communication
4.3 Grooming
4.4 Fighting
4.5 Hunting and feeding
4.6 Play
4.7 Reproduction
5 Ecology
5.1 Habitats
5.2 Feral cats
5.3 Impact on prey species
5.4 Impact on birds
6 Interaction with humans
6.1 Cat show
6.2 Cat café
6.3 Ailurophobia
6.4 Cat bites
6.5 Infections transmitted from cats to humans
6.6 History and mythology
6.6.1 Superstitions and cat burning
7 See also
8 Notes
9 References
10 External links
Taxonomy and evolution
Main article: Cat evolution
The domestic cat is believed to have evolved from the Near Eastern wildcat, whose range covers vast portions of the Middle East westward to the Atlantic coast of Africa.[22][23] Between 70,000 and 100,000 years ago the animal gave rise to the genetic lineage that eventually produced all domesticated cats,[24] having diverged from the Near Eastern wildcat around 8,000 BC in the Middle East.[12][15]
The felids are a rapidly evolving family of mammals that share a common ancestor only 10–15 million years ago[25] and include lions, tigers, cougars and many others. Within this family, domestic cats (Felis catus) are part of the genus Felis, which is a group of small cats containing about seven species (depending upon classification scheme).[1][26] Members of the genus are found worldwide and include the jungle cat (Felis chaus) of southeast Asia, European wildcat (F. silvestris silvestris), African wildcat (F. s. lybica), the Chinese mountain cat (F. bieti), and the Arabian sand cat (F. margarita), among others.[27]
The domestic cat was first classified as Felis catus by Carl Linnaeus in the 10th edition of his Systema Naturae published in 1758.[1][2] Because of modern phylogenetics, domestic cats are usually regarded as another subspecies of the wildcat, F. silvestris.[1][28][29] This has resulted in mixed usage of the terms, as the domestic cat can be called by its subspecies name, Felis silvestris catus.[1][28][29] Wildcats have also been referred to as various subspecies of F. catus,[29] but in 2003, the International Commission on Zoological Nomenclature fixed the name for wildcats as F. silvestris.[30] The most common name in use for the domestic cat remains F. catus. Sometimes, the domestic cat has been called Felis domesticus[31] as proposed by German naturalist J.C.P. Erxleben in 1777,[32] but these are not valid taxonomic names and have been used only rarely in scientific literature.[33] A population of Transcaucasian black feral cats was once classified as Felis daemon (Satunin 1904) but now this population is considered to be a part of domestic cat.[34]
All the cats in this genus share a common ancestor that is believed to have lived around 6–7 million years ago in the Near East (the Middle East).[35] The exact relationships within the Felidae are close but still uncertain,[36][37] e.g. the Chinese mountain cat is sometimes classified (under the name Felis silvestris bieti) as a subspecies of the wildcat, like the North African variety F. s. lybica.[28][36]
Ancient Egyptian sculpture of the cat goddess Bastet. The earliest evidence of felines as Egyptian deities comes from c. 3100 BC.
In comparison to dogs, cats have not undergone major changes during the domestication process, as the form and behavior of the domestic cat is not radically different from those of wildcats and domestic cats are perfectly capable of surviving in the wild.[38][39] Fully domesticated house cats often interbreed with feral F. catus populations,[40] producing hybrids such as the Kellas cat. This limited evolution during domestication means that hybridisation can occur with many other felids, notably the Asian leopard cat.[41] Several natural behaviors and characteristics of wildcats may have predisposed them for domestication as pets.[39] These traits include their small size, social nature, obvious body language, love of play and relatively high intelligence.[42]:12–17 Several small felid species may have an inborn tendency towards tameness.[39]
Cats have either a mutualistic or commensal relationship with humans. Two main theories are given about how cats were domesticated. In one, people deliberately tamed cats in a process of artificial selection as they were useful predators of vermin.[43] This has been criticized as implausible, because the reward for such an effort may have been too little; cats generally do not carry out commands and although they do eat rodents, other species such as ferrets or terriers may be better at controlling these pests.[28] The alternative idea is that cats were simply tolerated by people and gradually diverged from their wild relatives through natural selection, as they adapted to hunting the vermin found around humans in towns and villages.[28]
Nomenclature and etymology
The origin of the English word cat (Old English catt) and its counterparts in other Germanic languages (such as German Katze), descended from Proto-Germanic *kattōn-, is controversial. It has traditionally thought to be a borrowing from Late Latin cattus "domestic cat", from catta (used around 75 AD by Martial),[44][45] compare also Byzantine Greek κάττα, Portuguese and Spanish gato, French chat, Maltese qattus, Lithuanian katė, and Old Church Slavonic kotъ (kotka), among others.[46] The Late Latin word is generally thought to originate from an Afro-Asiatic language, but every proposed source word has presented problems. Many references refer to "Berber" (Kabyle) kaddîska "wildcat" and "Nubian kadīs" as possible sources or cognates, but M. Lionel Bender says the Nubian term is a loan from Arabic قِطَّة qiṭṭa.[47] Jean-Paul Savignac suggests the Latin word is from an Egyptian precursor of Coptic ϣⲁⲩ (šau) "tomcat" or its feminine form suffixed with -t,[48] but John Huehnergard says "the source [...] was clearly not Egyptian itself, where no analogous form is attested."[47] Huehnergard opines it is "equally likely that the forms might derive from an ancient Germanic word, imported into Latin and thence to Greek and to Syriac and Arabic". Guus Kroonen also considers the word to be native to Germanic (due to morphological alternations) and Northern Europe, and suggests that it might ultimately be borrowed from Uralic, cf. Northern Sami gađfe "female stoat" and Hungarian hölgy "stoat; lady, bride" from Proto-Uralic *käďwä "female (of a fur animal)".[49] In any case, cat is a classic case of a Wanderwort.
An alternative word is English puss (extended as pussy and pussycat). Attested only from the 16th century, it may have been introduced from Dutch poes or from Low German puuskatte, related to Swedish kattepus, or Norwegian pus, pusekatt. Similar forms exist in Lithuanian puižė and Irish puisín or puiscín. The etymology of this word is unknown, but it may have simply arisen from a sound used to attract a cat.[50][51]
A group of cats is referred to as a clowder or a glaring,[52] a male cat is called a tom or tomcat[53] (or a gib,[54] if neutered), an unaltered female is called a queen,[55] and a juvenile cat is referred to as a kitten. The male progenitor of a cat, especially a pedigreed cat, is its sire,[56] and its female progenitor is its dam.[57] In Early Modern English, the word kitten was interchangeable with the now obsolete word catling.[58]
A pedigreed cat is one whose ancestry is recorded by a cat fancier organization. A purebred cat is one whose ancestry contains only individuals of the same breed. Many pedigreed and especially purebred cats are exhibited as show cats. Cats of unrecorded, mixed ancestry are referred to as domestic short-haired or domestic long-haired cats, by coat type, or commonly as random-bred, moggies (chiefly British), or (using terms borrowed from dog breeding) mongrels or mutt-cats.
While the African wildcat is the ancestral subspecies from which domestic cats are descended, and wildcats and domestic cats can completely interbreed (being subspecies of the same species), several intermediate stages occur between domestic pet and pedigree cats on one hand and entirely wild animals on the other. The semiferal cat, a mostly outdoor cat, is not owned by any one individual, but is generally friendly to people and may be fed by several households. Feral cats are associated with human habitation areas and may be fed by people or forage for food, but are typically wary of human interaction.[40]
Biology
Anatomy
Main article: Cat anatomy
Diagram of the general anatomy of a male
Domestic cats are similar in size to the other members of the genus Felis, typically weighing between 4 and 5 kg (9 and 10 lb).[36] Some breeds, such as the Maine Coon, can occasionally exceed 11 kg (24 lb). Conversely, very small cats, less than 2 kg (4 lb), have been reported.[59] The world record for the largest cat is 21 kg (50 lb).[60] The smallest adult cat ever officially recorded weighed around 1 kg (2 lb).[60] Feral cats tend to be lighter as they have more limited access to food than house cats. In the Boston area, the average feral adult male will weigh 4 kg (9 lb) and average feral female 3 kg (7 lb).[61] Cats average about 23–25 cm (9–10 in) in height and 46 cm (18 in) in head/body length (males being larger than females), with tails averaging 30 cm (12 in) in length.[62]
Cats have seven cervical vertebrae, as do almost all mammals; 13 thoracic vertebrae (humans have 12); seven lumbar vertebrae (humans have five); three sacral vertebrae like most mammals (humans have five); and a variable number of caudal vertebrae in the tail (humans retain three to five caudal vertebrae, fused into an internal coccyx).[63]:11 The extra lumbar and thoracic vertebrae account for the cat's spinal mobility and flexibility. Attached to the spine are 13 ribs, the shoulder, and the pelvis.[63] :16 Unlike human arms, cat forelimbs are attached to the shoulder by free-floating clavicle bones which allow them to pass their body through any space into which they can fit their head.[64]
Cat skull
The cat skull is unusual among mammals in having very large eye sockets and a powerful and specialized jaw.[65]:35 Within the jaw, cats have teeth adapted for killing prey and tearing meat. When it overpowers its prey, a cat delivers a lethal neck bite with its two long canine teeth, inserting them between two of the prey's vertebrae and severing its spinal cord, causing irreversible paralysis and death.[66] Compared to other felines, domestic cats have narrowly spaced canine teeth, which is an adaptation to their preferred prey of small rodents, which have small vertebrae.[66] The premolar and first molar together compose the carnassial pair on each side of the mouth, which efficiently shears meat into small pieces, like a pair of scissors. These are vital in feeding, since cats' small molars cannot chew food effectively, and cats are largely incapable of mastication.[65]:37 Although cats tend to have better teeth than most humans, with decay generally less likely because of a thicker protective layer of enamel, a less damaging saliva, less retention of food particles between teeth, and a diet mostly devoid of sugar, they are nonetheless subject to occasional tooth loss and infection.[67]
Cats, like dogs, are digitigrades. They walk directly on their toes, with the bones of their feet making up the lower part of the visible leg.[68] Cats are capable of walking very precisely because, like all felines, they directly register; that is, they place each hind paw (almost) directly in the print of the corresponding fore paw, minimizing noise and visible tracks. This also provides sure footing for their hind paws when they navigate rough terrain. Unlike most mammals, when cats walk, they use a "pacing" gait; that is, they move the two legs on one side of the body before the legs on the other side. This trait is shared with camels and giraffes. As a walk speeds up into a trot, a cat's gait changes to be a "diagonal" gait, similar to that of most other mammals (and many other land animals, such as lizards): the diagonally opposite hind and fore legs move simultaneously.[69]
Like almost all members of the Felidae, cats have protractable and retractable claws.[70] In their normal, relaxed position, the claws are sheathed with the skin and fur around the paw's toe pads. This keeps the claws sharp by preventing wear from contact with the ground and allows the silent stalking of prey. The claws on the fore feet are typically sharper than those on the hind feet.[71] Cats can voluntarily extend their claws on one or more paws. They may extend their claws in hunting or self-defense, climbing, kneading, or for extra traction on soft surfaces. Most cats have five claws on their front paws, and four on their rear paws.[72] The fifth front claw (the dewclaw) is proximal to the other claws. More proximally is a protrusion which appears to be a sixth "finger". This special feature of the front paws, on the inside of the wrists, is the carpal pad, also found on the paws of big cats and dogs. It has no function in normal walking, but is thought to be an antiskidding device used while jumping. Some breeds of cats are prone to polydactyly (extra toes and claws).[72] These are particularly common along the northeast coast of North America.[73]
Physiology
Cats are familiar and easily kept animals, and their physiology has been particularly well studied; it generally resembles those of other carnivorous mammals, but displays several unusual features probably attributable to cats' descent from desert-dwelling species.[31] For instance, cats are able to tolerate quite high temperatures: Humans generally start to feel uncomfortable when their skin temperature passes about 38 °C (100 °F), but cats show no discomfort until their skin reaches around 52 °C (126 °F),[65]:46 and can tolerate temperatures of up to 56 °C (133 °F) if they have access to water.[74]
Normal physiological values[75]:330
Body temperature 38.6 °C (101.5 °F)
Heart rate 120–140 beats per minute
Breathing rate 16–40 breaths per minute
Thermograph of various body parts of a cat
Cats conserve heat by reducing the flow of blood to their skin and lose heat by evaporation through their mouths. Cats have minimal ability to sweat, with glands located primarily in their paw pads,[76] and pant for heat relief only at very high temperatures[77] (but may also pant when stressed). A cat's body temperature does not vary throughout the day; this is part of cats' general lack of circadian rhythms and may reflect their tendency to be active both during the day and at night.[78]:1 Cats' feces are comparatively dry and their urine is highly concentrated, both of which are adaptations to allow cats to retain as much water as possible.[31] Their kidneys are so efficient, they can survive on a diet consisting only of meat, with no additional water,[79] and can even rehydrate by drinking seawater.[80][78]:29While domestic cats are able to swim, they are generally reluctant to enter water as it quickly leads to exhaustion.[81]
Nutrition
Cats are obligate carnivores: their physiology has evolved to efficiently process meat, and they have difficulty digesting plant matter.[31] In contrast to omnivores such as rats, which only require about 4% protein in their diet, about 20% of a cat's diet must be protein.[31] A cat's gastrointestinal tract is adapted to meat eating, being much shorter than that of omnivores and having low levels of several of the digestive enzymes needed to digest carbohydrates.[82] These traits severely limit the cat's ability to digest and use plant-derived nutrients, as well as certain fatty acids.[82] Despite the cat's meat-oriented physiology, several vegetarian or vegan cat foods have been marketed that are supplemented with chemically synthesized taurine and other nutrients, in attempts to produce a complete diet. However, some of these products still fail to provide all the nutrients cats require,[83] and diets containing no animal products pose the risk of causing severe nutritional deficiencies.[84] However, veterinarians in the United States have expressed concern that many domestic cats are overfed.[85]
Cats do eat grass occasionally. A proposed explanation is that cats use grass as a source of folic acid. Another is that it is used to supply dietary fiber, helping the cat defecate more easily and expel parasites and other harmful material through feces and vomit.[86]
Cats are unusually dependent on a constant supply of the amino acid arginine, and a diet lacking arginine causes marked weight loss and can be rapidly fatal.[87] Arginine is an essential additive in cat food because cats have low levels of the enzymes aminotransferase and pyrroline-5-carboxylate which are responsible for the synthesis of ornithine and citrulline in the small intestine.[88] Citrulline would typically go on to the kidneys to make arginine, but because cats have a deficiency in the enzymes that make it, citrulline is not produced in adequate quantities to make arginine. Arginine is essential in the urea cycle in order to convert the toxic component ammonia into urea that can then be excreted in the urine. Because of its essential role, deficiency in arginine results in a build up of toxic ammonia and leads to hyperammonemia.[88] The symptoms of hyperammonemia include lethargy, vomiting, ataxia, hyperesthesia and can be serious enough to induce death and coma in a matter of days if a cat is being fed a arginine free diet. The quick onset of these symptoms is due to the fact that diets devoid in arginine will typically still contain all of the other amino acids, which will continue to be catabolized by the body producing mass amounts of ammonia that very quickly build up with no way of being excreted.
Another unusual feature is that the cat cannot produce taurine,[note 1] with a deficiency in this nutrient causing macular degeneration, wherein the cat's retina slowly breaks down, causing irreversible blindness.[31] This is due to the hepatic activity of cystinesulfinic acid decarboxylase being low in cats.[90] This limits the ability of cats to biosynthesize the taurine they need from its precursor, the amino acid cysteine, which ultimately results in inadequate taurine production needed for normal function.[90] Deficiencies in taurine result in compensated function of feline cardiovascular and reproductive systems.[90] These abnormalities can also be accompanied by developmental issues in the central nervous system along with degeneration of the retina.[90]
In order to produce the essential vitamin niacin for use in the cat, tryptophan is needed for conversion purposes. However, due to a competing pathway with acetyl coenzyme A (acetyl-CoA), niacin can become deficient and require supplementation.[91] This process occurs when an overactive enzyme, picolinic acid carboxylase, converts the vitamin B6 precursor picolinic acid into the alternate compound acetyl-CoA, instead of converting quinolinate into nictotinic acid mononlucleotide (niacin).[92] Niacin is required in cats as it supports enzyme function. If niacin is deficient in the diet, anorexia, weight loss and an increase in body temperature can result.[93]
Preformed vitamin A is required in the cat for retinal and reproductive health. Vitamin A is considered to be a fat-soluble vitamin and is seen as essential in a cat's diet. Normally, the conversion of beta-carotenes into vitamin A occurs in the intestine (more specifically the mucosal layer) of species, however cats lack the ability to undergo this process.[91] Both the kidney and liver are contributors to the use of vitamin A in the body of the majority of species while the cats liver does not produce the enzyme Beta-carotene 15,15'-monooxygenase which converts the beta-carotene into retinol (vitamin A).[94] To summarize: cats do not have high levels of this enzyme leading to the cleavage and oxidation of carotenoids not taking place.[92]
Vitamin D3 is a dietary requirement for cats as they lack the ability to synthesize vitamin D3 from sunlight.[95] Cats obtain high levels of the enzyme 7-dehydrocholestrol delta 7 reductase which causes immediate conversion of vitamin D3 from sunlight to 7-dehydrocholesterol.[96] This fat soluble vitamin is required in cats for bone formation through the promotion of calcium retention, along with nerve and muscle control through absorption of calcium and phosphorus.[96]
Cats, like all mammals, need to get linoleic acid, an essential fatty acid, from their diet. Most mammals can convert linoleic acid to arachidonic acid, as well as the omega 3 fatty acids (eicosapentaenoic acid and docosahexaenoic acid) through the activity of enzymes, but this process is very limited in cats.[91] The Δ6-desaturase enzyme eventually converts linoleic acid, which is in its salt form linoleate, to arachidonate (salt form of arachidonic acid) in the liver, but this enzyme has very little activity in cats.[91] This means that arachidonic acid is an essential fatty acid for cats as they lack the ability to create required amounts of linoleic acid. Deficiency of arachidonic acid in cats is related to problems in growth, can cause injury and inflammation to skin (e.g. around the mouth) decreased platelet aggregation, fatty liver, increase in birth defects of kittens whose queens were deficient during pregnancy, and reproductive failure in queens.[91] Arachidonic acid can also be metabolized to eicosanoids that create inflammatory responses which are needed to stimulate proper growth and repair mechanisms in the cat.[97]
The nutrient chart provides a list of the many nutrients cats require as well as the use of the nutrients in the body and the effects of the deficiency.
Senses
Main article: Cat senses
Reflection of camera flash from the tapetum lucidum
Cats have excellent night vision and can see at only one-sixth the light level required for human vision.[65]:43 This is partly the result of cat eyes having a tapetum lucidum, which reflects any light that passes through the retina back into the eye, thereby increasing the eye's sensitivity to dim light.[98] Another adaptation to dim light is the large pupils of cats' eyes. Unlike some big cats, such as tigers, domestic cats have slit pupils.[99] These slit pupils can focus bright light without chromatic aberration, and are needed since the domestic cat's pupils are much larger, relative to their eyes, than the pupils of the big cats.[99] At low light levels, a cat's pupils will expand to cover most of the exposed surface of its eyes.[100] However, domestic cats have rather poor color vision and (like most nonprimate mammals) have only two types of cones, optimized for sensitivity to blue and yellowish green; they have limited ability to distinguish between red and green.[101] A 1993 paper reported a response to middle wavelengths from a system other than the rods which might be due to a third type of cone. However, this appears to be an adaptation to low light levels rather than representing true trichromatic vision.[102]
Cats have excellent hearing and can detect an extremely broad range of frequencies. They can hear higher-pitched sounds than either dogs or humans, detecting frequencies from 55 Hz to 79,000 Hz, a range of 10.5 octaves, while humans and dogs both have ranges of about 9 octaves.[103][104] Cats can hear ultrasound, which is important in hunting[105] because many species of rodents make ultrasonic calls.[106] However, they do not communicate using ultrasound like rodents do. Cats' hearing is also sensitive and among the best of any mammal,[103] being most acute in the range of 500 Hz to 32 kHz.[107] This sensitivity is further enhanced by the cat's large movable outer ears (their pinnae), which both amplify sounds and help detect the direction of a noise.[105]
Cats have an acute sense of smell, due in part to their well-developed olfactory bulb and a large surface of olfactory mucosa, about 5.8 cm2 (0.90 in2) in area, which is about twice that of humans.[108] Cats are sensitive to pheromones such as 3-mercapto-3-methylbutan-1-ol,[109] which they use to communicate through urine spraying and marking with scent glands.[110] Many cats also respond strongly to plants that contain nepetalactone, especially catnip, as they can detect that substance at less than one part per billion.[111] About 70–80% of cats are affected by nepetalactone.[112] This response is also produced by other plants, such as silver vine (Actinidia polygama) and the herb valerian; it may be caused by the smell of these plants mimicking a pheromone and stimulating cats' social or sexual behaviors.[113]
Cats have relatively few taste buds compared to humans (470 or so versus more than 9,000 on the human tongue).[114] Domestic and wild cats share a gene mutation that keeps their sweet taste buds from binding to sugary molecules, leaving them with no ability to taste sweetness.[115] Their taste buds instead respond to amino acids, bitter tastes, and acids.[116] Cats and many other animals have a Jacobson's organ in their mouths that is used in the behavioral process of flehmening. It allows them to sense certain aromas in a way that humans cannot. Cats also have a distinct temperature preference for their food, preferring food with a temperature around 38 °C (100 °F) which is similar to that of a fresh kill and routinely rejecting food presented cold or refrigerated (which would signal to the cat that the "prey" item is long dead and therefore possibly toxic or decomposing).[114]
The whiskers of a cat are highly sensitive to touch.
To aid with navigation and sensation, cats have dozens of movable whiskers (vibrissae) over their body, especially their faces. These provide information on the width of gaps and on the location of objects in the dark, both by touching objects directly and by sensing air currents; they also trigger protective blink reflexes to protect the eyes from damage.[65]:47
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Comparison of cat righting reflexes in gravity versus zero gravity
Most breeds of cat have a noted fondness for settling in high places, or perching. In the wild, a higher place may serve as a concealed site from which to hunt; domestic cats may strike prey by pouncing from a perch such as a tree branch, as does a leopard.[117] Another possible explanation is that height gives the cat a better observation point, allowing it to survey its territory. During a fall from a high place, a cat can reflexively twist its body and right itself using its acute sense of balance and flexibility.[118] This is known as the cat righting reflex. An individual cat always rights itself in the same way, provided it has the time to do so, during a fall. The height required for this to occur is around 90 cm (3.0 ft). Cats without a tail (e.g. Manx cats) also have this ability, since a cat mostly moves its hind legs and relies on conservation of angular momentum to set up for landing, and the tail is little used for this feat.[119] An excellent sense of balance allows cats to move with great stability. A cat falling from heights of up to 3 meters can right itself and land on its paws.[120]
Health
Main articles: Cat health and Aging in cats
The average lifespan of pet cats has risen in recent years. In the early 1980s, it was about seven years,[121]:33[122] rising to 9.4 years in 1995[121]:33 and 12–15 years in 2014.[123][unreliable source?] However, cats have been reported as surviving into their 30s,[124] with the oldest known cat, Creme Puff, dying at a verified age of 38.[125]
Spaying or neutering increases life expectancy: one study found neutered male cats live twice as long as intact males, while spayed female cats live 62% longer than intact females.[121]:35 Having a cat neutered confers health benefits, because castrated males cannot develop testicular cancer, spayed females cannot develop uterine or ovarian cancer, and both have a reduced risk of mammary cancer.[126]
Despite widespread concern about the welfare of free-roaming cats, the lifespans of neutered feral cats in managed colonies compare favorably with those of pet cats.[127]:45[128]:1358 [129][130][131][132]
Diseases
Main article: Feline diseases
A wide range of health problems may affect cats, including infectious diseases, parasites, injuries, and chronic disease. Vaccinations are available for many of these diseases, and domestic cats are regularly given treatments to eliminate parasites such as worms and fleas.[133]
Genetics
Main article: Cat genetics
The domesticated cat and its closest wild ancestor are both diploid organisms that possess 38 chromosomes[134] and roughly 20,000 genes.[135] About 250 heritable genetic disorders have been identified in cats, many similar to human inborn errors.[136] The high level of similarity among the metabolism of mammals allows many of these feline diseases to be diagnosed using genetic tests that were originally developed for use in humans, as well as the use of cats as animal models in the study of the human diseases.[137][138]
Behavior
See also: Cat behavior and Cat intelligence
A cat on a fence
Outdoor cats are active both day and night, although they tend to be slightly more active at night.[139][140] The timing of cats' activity is quite flexible and varied, which means house cats may be more active in the morning and evening, as a response to greater human activity at these times.[141] Although they spend the majority of their time in the vicinity of their home, housecats can range many hundreds of meters from this central point, and are known to establish territories that vary considerably in size, in one study ranging from 7 to 28 hectares (17–69 acres).[140]
Cats conserve energy by sleeping more than most animals, especially as they grow older. The daily duration of sleep varies, usually between 12 and 16 hours, with 13 and 14 being the average. Some cats can sleep as much as 20 hours. The term "cat nap" for a short rest refers to the cat's tendency to fall asleep (lightly) for a brief period. While asleep, cats experience short periods of rapid eye movement sleep often accompanied by muscle twitches, which suggests they are dreaming.[142]
Sociability
Social grooming
Although wildcats are solitary, the social behavior of domestic cats is much more variable and ranges from widely dispersed individuals to feral cat colonies that gather around a food source, based on groups of co-operating females.[143][144] Within such groups, one cat is usually dominant over the others.[33] Each cat in a colony holds a distinct territory, with sexually active males having the largest territories, which are about 10 times larger than those of female cats and may overlap with several females' territories.[110] These territories are marked by urine spraying, by rubbing objects at head height with secretions from facial glands, and by defecation.[110] Between these territories are neutral areas where cats watch and greet one another without territorial conflicts. Outside these neutral areas, territory holders usually chase away stranger cats, at first by staring, hissing, and growling, and if that does not work, by short but noisy and violent attacks. Despite some cats cohabiting in colonies, they do not have a social survival strategy, or a pack mentality, and always hunt alone.[145]
Cat with an Alaskan Malamute dog
However, some pet cats are poorly socialized. In particular, older cats may show aggressiveness towards newly arrived kittens, which may include biting and scratching; this type of behavior is known as feline asocial aggression.[146]
Though cats and dogs are often characterized as natural enemies, they can live together if correctly socialized.[147]
Life in proximity to humans and other domestic animals has led to a symbiotic social adaptation in cats, and cats may express great affection toward humans or other animals. Ethologically, the human keeper of a cat may function as a sort of surrogate for the cat's mother,[148] and adult housecats live their lives in a kind of extended kittenhood,[149] a form of behavioral neoteny. The high-pitched sounds housecats make to solicit food may mimic the cries of a hungry human infant, making them particularly hard for humans to ignore.[150]
Domestic cat's scent rubbing behavior towards humans or other cats is thought to be a feline means for social bonding.[151]
Communication
Main article: Cat communication
Domestic cats use many vocalizations for communication, including purring, trilling, hissing, growling/snarling, grunting, and several different forms of meowing.[7] (By contrast, feral cats are generally silent.)[152]:208 Their types of body language, including position of ears and tail, relaxation of the whole body, and kneading of the paws, are all indicators of mood. The tail and ears are particularly important social signal mechanisms in cats;[153][154] for example, a raised tail acts as a friendly greeting, and flattened ears indicates hostility. Tail-raising also indicates the cat's position in the group's social hierarchy, with dominant individuals raising their tails less often than subordinate animals.[154] Nose-to-nose touching is also a common greeting and may be followed by social grooming, which is solicited by one of the cats raising and tilting its head.[144]
Purring may have developed as an evolutionary advantage as a signalling mechanism of reassurance between mother cats and nursing kittens. Post-nursing cats often purr as a sign of contentment: when being petted, becoming relaxed,[155][156] or eating. The mechanism by which cats purr is elusive. The cat has no unique anatomical feature that is clearly responsible for the sound.[157] It was, until recent times, believed that only the cats of the Felis genus could purr. However, felids of the genus Panthera (tiger, lion, jaguar, and leopard) also produce sounds similar to purring, but only when exhaling.[158]
Grooming
The hooked papillae on a cat's tongue act like a hairbrush to help clean and detangle fur.
Play media
A tabby housecat uses its brush-like tongue to groom itself, licking its fur to straighten it.
Cats are known for spending considerable amounts of time licking their coat to keep it clean.[159] The cat's tongue has backwards-facing spines about 500 μm long, which are called papillae. These contain keratin which makes them rigid[160] so the papillae act like a hairbrush. Some cats, particularly longhaired cats, occasionally regurgitate hairballs of fur that have collected in their stomachs from grooming. These clumps of fur are usually sausage-shaped and about 2–3 cm (0.8–1.2 in) long. Hairballs can be prevented with remedies that ease elimination of the hair through the gut, as well as regular grooming of the coat with a comb or stiff brush.[159]
Fighting
Among domestic cats, males are more likely to fight than females.[161] Among feral cats, the most common reason for cat fighting is competition between two males to mate with a female. In such cases, most fights are won by the heavier male.[162] Another common reason for fighting in domestic cats is the difficulty of establishing territories within a small home.[161] Female cats also fight over territory or to defend their kittens. Neutering will decrease or eliminate this behavior in many cases, suggesting that the behavior is linked to sex hormones.[163]
An arched back, raised fur and an open-mouthed hiss can all be signs of aggression in a domestic cat.
When cats become aggressive, they try to make themselves appear larger and more threatening by raising their fur, arching their backs, turning sideways and hissing or spitting.[153] Often, the ears are pointed down and back to avoid damage to the inner ear and potentially listen for any changes behind them while focused forward. They may also vocalize loudly and bare their teeth in an effort to further intimidate their opponent. Fights usually consist of grappling and delivering powerful slaps to the face and body with the forepaws as well as bites. Cats also throw themselves to the ground in a defensive posture to rake their opponent's belly with their powerful hind legs.[164]
Serious damage is rare, as the fights are usually short in duration, with the loser running away with little more than a few scratches to the face and ears. However, fights for mating rights are typically more severe and injuries may include deep puncture wounds and lacerations. Normally, serious injuries from fighting are limited to infections of scratches and bites, though these can occasionally kill cats if untreated. In addition, bites are probably the main route of transmission of feline immunodeficiency virus.[165] Sexually active males are usually involved in many fights during their lives, and often have decidedly battered faces with obvious scars and cuts to their ears and nose.[166]
Hunting and feeding
A cat that is playing with a caught mouse. Cats play with their prey to weaken or exhaust them before making a kill.
A domestic cat with its prey
Cats hunt small prey, primarily birds and rodents,[167] and are often used as a form of pest control.[168][169] Domestic cats are a major predator of wildlife in the United States, killing an estimated 1.4–3.7 billion birds and 6.9–20.7 billion mammals annually.[170][171] The bulk of predation in the United States is done by 80 million feral and stray cats. Effective measures to reduce this population are elusive, meeting opposition from cat enthusiasts.[170][171] In the case of free-ranging pets, equipping cats with bells and not letting them out at night will reduce wildlife predation.[167]
Free-fed feral cats and house cats tend to consume many small meals in a single day, although the frequency and size of meals varies between individuals.[145] Cats use two hunting strategies, either stalking prey actively, or waiting in ambush until an animal comes close enough to be captured.[172] Although it is not certain, the strategy used may depend on the prey species in the area, with cats waiting in ambush outside burrows, but tending to actively stalk birds.[173]:153
Perhaps the best known element of cats' hunting behavior, which is commonly misunderstood and often appalls cat owners because it looks like torture, is that cats often appear to "play" with prey by releasing it after capture. This behavior is due to an instinctive imperative to ensure that the prey is weak enough to be killed without endangering the cat.[174] This behavior is referred to in the idiom "cat-and-mouse game" or simply "cat and mouse".
Another poorly understood element of cat hunting behavior is the presentation of prey to human guardians. Ethologist Paul Leyhausen proposed that cats adopt humans into their social group and share excess kill with others in the group according to the dominance hierarchy, in which humans are reacted to as if they are at, or near, the top.[175] Anthropologist and zoologist Desmond Morris, in his 1986 book Catwatching, suggests, when cats bring home mice or birds, they are attempting to teach their human to hunt, or trying to help their human as if feeding "an elderly cat, or an inept kitten".[176][177] Morris's hypothesis is inconsistent with the fact that male cats also bring home prey, despite males having negligible involvement with raising kittens.[173]:153
Domestic cats select food based on its temperature, smell and texture; they dislike chilled foods and respond most strongly to moist foods rich in amino acids, which are similar to meat.[84][145] Cats may reject novel flavors (a response termed neophobia) and learn quickly to avoid foods that have tasted unpleasant in the past.[145] They may also avoid sugary foods and milk. Most adult cats are lactose intolerant; the sugars in milk are not easily digested and may cause soft stools or diarrhea.[145][178] They can also develop odd eating habits. Some cats like to eat or chew on other things, most commonly wool, but also plastic, cables, paper, string, aluminum foil, or even coal. This condition, pica, can threaten their health, depending on the amount and toxicity of the items eaten.[179][180]
Though cats usually prey on animals less than half their size, a feral cat in Australia has been photographed killing an adult pademelon of around the cat's weight at 4 kg (8.8 lb).[181]
Since cats lack lips[182] to create suction, they use a lapping method with the tongue to draw liquid upwards into their mouths. Lapping at a rate of four times a second, the cat touches the smooth tip of its tongue to the surface of the water, and quickly retracts it like a corkscrew, drawing water upwards.[183]
Play
Main article: Cat play and toys
Play media
Play fight between kittens, age 14 weeks
Domestic cats, especially young kittens, are known for their love of play. This behavior mimics hunting and is important in helping kittens learn to stalk, capture, and kill prey.[184] Cats also engage in play fighting, with each other and with humans. This behavior may be a way for cats to practice the skills needed for real combat, and might also reduce any fear they associate with launching attacks on other animals.[185]
Owing to the close similarity between play and hunting, cats prefer to play with objects that resemble prey, such as small furry toys that move rapidly, but rapidly lose interest (they become habituated) in a toy they have played with before.[186] Cats also tend to play with toys more when they are hungry.[187] String is often used as a toy, but if it is eaten, it can become caught at the base of the cat's tongue and then move into the intestines, a medical emergency which can cause serious illness, even death.[188] Owing to the risks posed by cats eating string, it is sometimes replaced with a laser pointer's dot, which cats may chase.[189]
Reproduction
See also: Kitten
When cats mate, the tomcat (male) bites the scruff of the female's neck as she assumes a position conducive to mating known as lordosis behavior.
Radiography of a pregnant cat (about one month and a half)
Female cats are seasonally polyestrous, which means they may have many periods of heat over the course of a year, the season beginning in spring and ending in late autumn. Heat periods occur about every two weeks and last about 4 to 7 days.[190] Multiple males will be attracted to a female in heat. The males will fight over her, and the victor wins the right to mate. At first, the female rejects the male, but eventually the female allows the male to mate. The female utters a loud yowl as the male pulls out of her because a male cat's penis has a band of about 120–150 backwards-pointing penile spines, which are about 1 mm long; upon withdrawal of the penis, the spines rake the walls of the female's vagina, which acts to induce ovulation. This act also occurs to clear the vagina of other sperm in the context of a second (or more) mating, thus giving the later males a larger chance of conception.[191]
After mating, the female washes her vulva thoroughly. If a male attempts to mate with her at this point, the female will attack him. After about 20 to 30 minutes, once the female is finished grooming, the cycle will repeat.[190]
Because ovulation is not always triggered by a single mating, females may not be impregnated by the first male with which they mate.[192] Furthermore, cats are superfecund; that is, a female may mate with more than one male when she is in heat, with the result that different kittens in a litter may have different fathers.[190]
A newborn kitten
At 124 hours after conception, the morula forms. At 148 hours, early blastocysts form. At 10–12 days, implantation occurs.[193][194]
The gestation period for cats is between 64 and 67 days, with an average of 66 days.[195] The size of a litter usually is three to five kittens, with the first litter usually smaller than subsequent litters. Kittens are weaned between six and seven weeks old, and cats normally reach sexual maturity at 5–10 months (females) and to 5–7 months (males), although this can vary depending on breed.[190] Females can have two to three litters per year, so may produce up to 150 kittens in their breeding span of around ten years.[190]
Cats are ready to go to new homes at about 12 weeks of age,[196] when they are ready to leave their mother. They can be surgically sterilized (spayed or castrated) as early as 7 weeks to limit unwanted reproduction.[197] This surgery also prevents undesirable sex-related behavior, such as aggression, territory marking (spraying urine) in males and yowling (calling) in females. Traditionally, this surgery was performed at around six to nine months of age, but it is increasingly being performed before puberty, at about three to six months.[198] In the US, about 80% of household cats are neutered.[199]
Ecology
Habitats
A cat in snowy weather
Cats are a cosmopolitan species and are found across much of the world.[38] Geneticist Stephen James O'Brien, of the National Cancer Institute in Frederick, Maryland, remarked on how successful cats have been in evolutionary terms: "Cats are one of evolution's most charismatic creatures. They can live on the highest mountains and in the hottest deserts."[200] They are extremely adaptable and are now present on all continents except Antarctica, and on 118 of the 131 main groups of islands—even on isolated islands such as the Kerguelen Islands.[201][202]
Feral cats can live in forests, grasslands, tundra, coastal areas, agricultural land, scrublands, urban areas, and wetlands.[203] Their habitats even include small oceanic islands with no human inhabitants.[204] Further, the close relatives of domestic cats, the African wildcat (Felis silvestris lybica) and the Arabian sand cat (Felis margarita) both inhabit desert environments,[28] and domestic cats still show similar adaptations and behaviors.[31] The cat's ability to thrive in almost any terrestrial habitat has led to its designation as one of the world's worst invasive species.[205]
As domestic cats are little altered from wildcats, they can readily interbreed. This hybridization poses a danger to the genetic distinctiveness of some wildcat populations, particularly in Scotland and Hungary and possibly also the Iberian Peninsula.[41]
Feral cats
Main article: Feral cat
Feral farm cat
Feral cats are domestic cats that were born in or have reverted to a wild state. They are unfamiliar with and wary of humans and roam freely in urban and rural areas.[9] The numbers of feral cats is not known, but estimates of the US feral population range from 25 to 60 million.[9] Feral cats may live alone, but most are found in large colonies, which occupy a specific territory and are usually associated with a source of food.[206] Famous feral cat colonies are found in Rome around the Colosseum and Forum Romanum, with cats at some of these sites being fed and given medical attention by volunteers.[207]
Public attitudes towards feral cats vary widely, ranging from seeing them as free-ranging pets, to regarding them as vermin.[208] One common approach to reducing the feral cat population is termed 'trap-neuter-return', where the cats are trapped, neutered, immunized against diseases such as rabies and the feline Panleukopenia and Leukemia viruses, and then released.[209] Before releasing them back into their feral colonies, the attending veterinarian often nips the tip off one ear to mark it as neutered and inoculated, since these cats may be trapped again. Volunteers continue to feed and give care to these cats throughout their lives. Given this support, their lifespans are increased, and behavior and nuisance problems caused by competition for food are reduced.[206]
Impact on prey species
Carrying half of a rabbit
To date, little scientific data is available to assess the impact of cat predation on prey populations outside of agricultural situations. Even well-fed domestic cats may hunt and kill, mainly catching small mammals, but also birds, amphibians, reptiles, fish, and invertebrates.[167][210] Hunting by domestic cats may be contributing to the decline in the numbers of birds in urban areas, although the importance of this effect remains controversial.[211] In the wild, the introduction of feral cats during human settlement can threaten native species with extinction.[204] In many cases, controlling or eliminating the populations of non-native cats can produce a rapid recovery in native animals.[212] However, the ecological role of introduced cats can be more complicated. For example, cats can control the numbers of rats, which also prey on birds' eggs and young, so a cat population can protect an endangered bird species by suppressing mesopredators.[213]
In isolated landmasses, such as Australasia, there are often no other native, medium-sized quadrupedal predators (including other feline species); this tends to exacerbate the impact of feral cats on small native animals.[214] Native species such as the New Zealand kakapo and the Australian bettong, for example, tend to be more ecologically vulnerable and behaviorally "naive", when faced with predation by cats.[215] Feral cats have had a major impact on these native species and have played a leading role in the endangerment and extinction of many animals.[216]
Even in places with ancient and numerous cat populations, such as Western Europe, cats appear to be growing in number and independently of their environments' carrying capacity (such as the numbers of prey available).[217][218] This may be explained, at least in part, by an abundance of food, from sources including feeding by pet owners and scavenging. For instance, research in Britain suggests that a high proportion of cats hunt only "recreationally"[218], and in South Sweden, where research in 1982 found that the population density of cats was as high as 2,000 per square kilometre (5,200/sq mi).[217]
In agricultural settings, cats can be effective at keeping mouse and rat populations low, but only if rodent harborage locations are kept under control.[219][220] While cats are effective at preventing rodent population explosions, they are not effective for eliminating pre-existing severe infestations.[221]
Impact on birds
A black cat eating a house sparrow
The domestic cat is a significant predator of birds. UK assessments indicate they may be accountable for an estimated 64.8 million bird deaths each year.[167] A 2012 study suggests feral cats may kill several billion birds each year in the United States.[222] Certain species appear more susceptible than others; for example, 30% of house sparrow mortality is linked to the domestic cat.[223] In the recovery of ringed robins (Erithacus rubecula) and dunnocks (Prunella modularis), 31% of deaths were a result of cat predation.[224] In parts of North America, the presence of larger carnivores such as coyotes which prey on cats and other small predators reduces the effect of predation by cats and other small predators such as opossums and raccoons on bird numbers and variety.[225] The proposal that cat populations will increase when the numbers of these top predators decline is called the mesopredator release hypothesis.
On islands, birds can contribute as much as 60% of a cat's diet.[226] In nearly all cases, however, the cat cannot be identified as the sole cause for reducing the numbers of island birds, and in some instances, eradication of cats has caused a 'mesopredator release' effect;[227] where the suppression of top carnivores creates an abundance of smaller predators that cause a severe decline in their shared prey. Domestic cats are, however, known to be a contributing factor to the decline of many species, a factor that has ultimately led, in some cases, to extinction. The South Island piopio, Chatham rail,[224] the New Zealand merganser,[228] and the common diving petrel[229] are a few from a long list, with the most extreme case being the flightless Lyall's wren, which was driven to extinction only a few years after its discovery.[230][231]
Some of the same factors that have promoted adaptive radiation of island avifauna over evolutionary time appear to promote vulnerability to non-native species in modern time. The susceptibility of many island birds is undoubtedly due to evolution in the absence of mainland predators, competitors, diseases, and parasites, in addition to lower reproductive rates and extended incubation periods.[232] The loss of flight, or reduced flying ability is also characteristic of many island endemics.[233] These biological aspects have increased vulnerability to extinction in the presence of introduced species, such as the domestic cat.[234] Equally, behavioral traits exhibited by island species, such as "predatory naivety"[235] and ground-nesting,[232] have also contributed to their susceptibility.
Interaction with humans
Main article: Human interaction with cats
Cats and people
Cats are common pets throughout the world, and their worldwide population exceeds 500 million.[12] Although cat guardianship has commonly been associated with women,[236] a 2007 Gallup poll reported that men and women in the United States of America were equally likely to own a cat.[237]
As well as being kept as pets, cats are also used in the international fur[238] and leather industries for making coats, hats, blankets and stuffed toys;[239] and shoes, gloves and musical instruments respectively[240] (about 24 cats are needed to make a cat fur coat).[241] This use has now been outlawed in the United States, Australia, and the European Union.[242] Cat pelts have been used for superstitious purposes as part of the practise of witchcraft,[243] and are still made into blankets in Switzerland as folk remedies believed to help rheumatism.[244] In the Western intellectual tradition, the idea of cats as everyday objects have served to illustrate problems of quantum mechanics in the Schrödinger's cat thought experiment.
A few attempts to build a cat census have been made over the years, both through associations or national and international organizations (such as the Canadian Federation of Humane Societies's one[245]) and over the Internet,[246][247] but such a task does not seem simple to achieve. General estimates for the global population of domestic cats range widely from anywhere between 200 million to 600 million.[248][249][250][251][252][253]
Cat show
Main article: Cat show
A cat show is a judged event where the owners of cats compete to win titles in various cat registering organizations by entering their cats to be judged after a breed standard. Both pedigreed and companion (or moggy) cats are admissible, although the rules differ from organization to organization. Cats are compared to a breed standard, and the owners of those judged to be closest to it are awarded a prize. Moggies are judged based on their temperament. Often, at the end of the year, all of the points accrued at various shows are added up and more national and regional titles are awarded.
Cat café
Main article: Cat café
A cat café is a theme café whose attraction is cats that can be watched and played with. Patrons pay a cover fee, generally hourly and thus cat cafés can be seen as a form of supervised indoor pet rental.
Ailurophobia
Main article: Ailurophobia
Ailurophobia is a human phobia of cats; however, the term is often associated with humans that have a hatred of cats.[254]
Cat bites
Main article: Cat bite
Cats may bite humans when provoked, during play or when aggressive. Complications from cat bites can develop.[255] A cat bite differs from the bites of other pets. This is because the teeth of a cat are sharp and pointed causing deep punctures. Skin usually closes rapidly over the bite and traps microorganisms that cause infection.[256][255]
Infections transmitted from cats to humans
Main article: Feline zoonosis
Cats can be infected or infested with viruses, bacteria, fungus, protozoans, arthropods or worms that can transmit diseases to humans.[257] In some cases, the cat exhibits no symptoms of the disease,[258] However, the same disease can then become evident in a human. The likelihood that a person will become diseased depends on the age and immune status of the person. Humans who have cats living in their home or in close association are more likely to become infected, however, those who do not keep cats as pets might also acquire infections from cat feces and parasites exiting the cat's body.[257][259] Some of the infections of most concern include salmonella, cat scratch disease and toxoplasmosis.[258]
History and mythology
Main articles: Cultural depictions of cats and Cats in ancient Egypt
The ancient Egyptians mummified dead cats out of respect in the same way that they mummified people.[260]
Ancient Roman mosaic of a cat killing a partridge from the House of the Faun in Pompeii
A 19th-century drawing of a tabby cat
Traditionally, historians tended to think ancient Egypt was the site of cat domestication, owing to the clear depictions of house cats in Egyptian paintings about 3,600 years old.[28] However, in 2004, a Neolithic grave excavated in Shillourokambos, Cyprus, contained the skeletons, laid close to one another, of both a human and a cat. The grave is estimated to be 9,500 years old, pushing back the earliest known feline–human association significantly.[15][261][262] The cat specimen is large and closely resembles the African wildcat, rather than present-day domestic cats. This discovery, combined with genetic studies, suggests cats were probably domesticated in the Middle East, in the Fertile Crescent around the time of the development of agriculture, and then were brought to Cyprus and Egypt.[14][19] Direct evidence for the domestication of cats 5,300 years ago in Quanhucun, China has been published by archaeologists and paleontologists from the University of Washington and Chinese Academy of Sciences. The cats are believed to have been attracted to the village by rodents, which in turn were attracted by grain cultivated and stored by humans.[263]
In ancient Egypt, cats were sacred animals, with the goddess Bastet often depicted in cat form, sometimes taking on the war-like aspect of a lioness.[264]:220 Killing a cat was absolutely forbidden[260] and the Greek historian Herodotus reports that, whenever a household cat died, the entire family would mourn and shave their eyebrows.[260] Families took their dead cats to the sacred city of Bubastis,[260] where they were embalmed and buried in sacred repositories.[260] Domestic cats were probably first introduced to Greece and southern Italy in the fifth century BC by the Phoenicians.[265] The earliest unmistakable evidence of the Greeks having domestic cats comes from two coins from Magna Graecia dating to the mid-fifth century BC showing Iokastos and Phalanthos, the legendary founders of Rhegion and Taras respectively, playing with their pet cats.[266]:57–58[267]
Housecats seem to have been extremely rare among the ancient Greeks and Romans;[267] Herodotus expressed astonishment at the domestic cats in Egypt, because he had only ever seen wildcats.[267] Even during later times, weasels were far more commonly kept as pets[267] and weasels, not cats, were seen as the ideal rodent-killers.[267] The usual ancient Greek word for "cat" was ailouros, meaning "thing with the waving tail",[266]:57[267] but this word could also be applied to any of the "various long-tailed carnivores kept for catching mice".[267] Cats are rarely mentioned in ancient Greek literature,[267] but Aristotle does remark in his History of Animals that "female cats are naturally lecherous."[266]:74[267] The Greeks later syncretized their own goddess Artemis with the Egyptian goddess Bastet, adopting Bastet's associations with cats and ascribing them to Artemis.[266]:77–79 In Ovid's Metamorphoses, when the gods flee to Egypt and take animal forms, the goddess Diana (the Roman equivalent of Artemis) turns into a cat.[266]:79 Cats eventually displaced ferrets as the pest control of choice because they were more pleasant to have around the house and were more enthusiastic hunters of mice.[268] During the Middle Ages, many of Artemis's associations with cats were grafted onto the Virgin Mary.[268] Cats are often shown in icons of Annunciation and of the Holy Family[268] and, according to Italian folklore, on the same night that Mary gave birth to Jesus, a virgin cat in Bethlehem gave birth to a kitten.[268] Domestic cats were spread throughout much of the rest of the world during the Age of Discovery, as ships' cats were carried on sailing ships to control shipboard rodents and as good-luck charms.[264]:223
Several ancient religions believed cats are exalted souls, companions or guides for humans, that are all-knowing but mute so they cannot influence decisions made by humans. In Japan, the maneki neko cat is a symbol of good fortune.[269] In Norse mythology, Freyja, the goddess of love, beauty, and fertility, is depicted as riding a chariot drawn by cats.[270] In Jewish legend, the first cat was living in the house of the first man Adam as a pet that get rid of mice from the house.[271] The cat was once partnering with the first dog before the latter broke an oath they had made which resulted in enmity between the descendants of these two animals.[271] It is also written that neither cats nor foxes are represented in the water, while every other animal has an incarnation species in the water.[271] Although no species are sacred in Islam, cats are revered by Muslims. Some Western writers have stated Muhammad had a favorite cat, Muezza.[272] He is reported to have loved cats so much, "he would do without his cloak rather than disturb one that was sleeping on it".[273] The story has no origin in early Muslim writers, and seems to confuse a story of a later Sufi saint, Ahmed ar-Rifa'i, centuries after Muhammad.[274] One of the companions of Muhammad was known as "Abu Hurayrah" (Father of the Kitten), in reference to his documented affection to cats.[275]
Superstitions and cat burning
Some cultures are superstitious about black cats, ascribing either good or bad luck to them.
Many cultures have negative superstitions about cats. An example would be the belief that a black cat "crossing one's path" leads to bad luck, or that cats are witches' familiars used to augment a witch's powers and skills. The killing of cats in Medieval Ypres, Belgium, is commemorated in the innocuous present-day Kattenstoet (cat parade).[276] In medieval France, cats would be burnt alive as a form of entertainment. According to Norman Davies, the assembled people "shrieked with laughter as the animals, howling with pain, were singed, roasted, and finally carbonized".[277]
"It was the custom to burn a basket, barrel, or sack full of live cats, which was hung from a tall mast in the midst of the bonfire; sometimes a fox was burned. The people collected the embers and ashes of the fire and took them home, believing that they brought good luck. The French kings often witnessed these spectacles and even lit the bonfire with their own hands. In 1648 Louis XIV, crowned with a wreath of roses and carrying a bunch of roses in his hand, kindled the fire, danced at it and partook of the banquet afterwards in the town hall. But this was the last occasion when a monarch presided at the midsummer bonfire in Paris. At Metz midsummer fires were lighted with great pomp on the esplanade, and a dozen cats, enclosed in wicker cages, were burned alive in them, to the amusement of the people. Similarly at Gap, in the department of the Hautes-Alpes, cats used to be roasted over the midsummer bonfire."[278]
According to a myth in many cultures, cats have multiple lives. In many countries, they are believed to have nine lives, but in Italy, Germany, Greece, Brazil and some Spanish-speaking regions, they are said to have seven lives,[279][280] while in Turkish and Arabic traditions, the number of lives is six.[281] The myth is attributed to the natural suppleness and swiftness cats exhibit to escape life-threatening situations. Also lending credence to this myth is the fact that falling cats often land on their feet, using an instinctive righting reflex to twist their bodies around. Nonetheless, cats can still be injured or killed by a high fall.[282]
See also
Book: Cat
Cats portal
Mammals portal
Aging in cats
Animal testing on cats
Animal track
Cancer in cats
Cat and mouse (cat-and-mouse game)
Cat burning
Cat intelligence
Cat lady
Cats and the Internet
Dog–cat relationship
Dried cat
List of cat breeds
List of cat documentaries
List of cats
List of fictional cats and felines
Pet door including cat flap
Pet first aid
Popular cat names
Trap-neuter-return
Cats by location
Cats in ancient Egypt
Cats in Australia
Cats in New Zealand
Cats in the United States
Notes
Jump up ^ Taurine is sometimes called an amino acid, and indeed is an acid containing an amino group, it is not an amino acid in the usual biochemical meaning of the term, which refers to compounds containing both an amino and a carboxyl group.[89]
References
^ Jump up to: a b c d e f Wozencraft, W.C. (2005). "Species Felis catus". In Wilson, D.E.; Reeder, D.M. Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Johns Hopkins University Press. pp. 534–535. ISBN 978-0-8018-8221-0. OCLC 62265494.
^ Jump up to: a b Linnaeus, C. (1758). Systema naturae (in Latin). 1 (10th ed.). Stockholm: Lars Salvius. p. 42. Archived from the original on 31 August 2017. Retrieved 4 September 2017.
Jump up ^ See Opinion 2027
Jump up ^ "ITIS Standard Report Page: Felis catus domestica". ITIS Online Database. Archived from the original on 19 January 2012. Retrieved 14 December 2011.
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Jump up ^ Bartleby: Ailurophobes and Other Cat-Haters Archived 23 December 2017 at the Wayback Machine.
^ Jump up to: a b "First Aid: Animal Bites". The Nemours Foundation. 2017. Archived from the original on 18 May 2017. Retrieved 18 June 2017.
Jump up ^ "Animal Bites". Handcare.org. American Society for Surgery of the Hand. 2017. Archived from the original on 28 July 2017. Retrieved 18 June 2017.
^ Jump up to: a b Chomel, Bruno (2014). "Emerging and Re-Emerging Zoonoses of Dogs and Cats". Animals. 4 (3): 434–445. doi:10.3390/ani4030434. ISSN 2076-2615. PMC 4494318  . PMID 26480316.
^ Jump up to: a b "Cats". Ohio Department of Health. 21 January 2015. Archived from the original on 27 November 2016. Retrieved 26 November 2016.
Jump up ^ Stull, J. W.; Brophy, J.; Weese, J. S. (2015). "Reducing the risk of pet-associated zoonotic infections". Canadian Medical Association Journal. 187 (10): 736–743. doi:10.1503/cmaj.141020. ISSN 0820-3946. PMC 4500695  . PMID 25897046.
^ Jump up to: a b c d e Clutton-Brock, Juliet (1999) [1987]. A Natural History of Domesticated Mammals (Second ed.). Cambridge, England: Cambridge University Press. p. 138. ISBN 0-521-63495-4.
Jump up ^ Muir, Hazel (8 April 2004). "Ancient Remains Could Be Oldest Pet Cat". New Scientist. Archived from the original on 21 December 2007. Retrieved 23 November 2007.
Jump up ^ Walton, Marsha (9 April 2004). "Ancient Burial Looks Like Human and Pet Cat". CNN. Archived from the original on 22 December 2007. Retrieved 23 November 2007.
Jump up ^ "Gatos fueron domesticados en China hace 5.300 años". La Nación (in Spanish). Archived from the original on 1 July 2014. Retrieved 9 September 2014.
^ Jump up to: a b Mason, I. L. (1984). Evolution of Domesticated Animals. Prentice Hall Press. ISBN 0-582-46046-8.
Jump up ^ Mark, Joshua J. (17 February 2012). "Cats in the Ancient World". Ancient History Encyclopedia. ancient.eu. Archived from the original on 2 January 2018.
^ Jump up to: a b c d e Engels, Donald W. (2001) [1999]. Classical Cats: The Rise and Fall of the Sacred Cat. New York City, New York and London, England: Routledge. ISBN 0-415-26162-7.
^ Jump up to: a b c d e f g h i Rogers, Katherine M. (2006). Cat. London, England: Reaktion Books. pp. 18–20. ISBN 978-1-86189-292-8.
^ Jump up to: a b c d Beadle, Muriel (1977). Cat. New York City, New York: Simon & Schuster. p. 76. ISBN 978-0671224516.
Jump up ^ Pate, Alan (2008). "Maneki Neko: Feline Fact & Fiction". Daruma Magazine. Amagasaki, Japan: Takeguchi Momoko. Archived from the original on 30 December 2012. Retrieved 30 December 2012.
Jump up ^ Faulkes, Anthony (1995). Edda. p. 24. ISBN 0-460-87616-3.
^ Jump up to: a b c Ginzberg, Louis (1909). The Legends of the Jews Vol. I : The Sixth Day (Translated by Henrietta Szold) Philadelphia: Jewish Publication Society
Jump up ^ Geyer, Georgie Anne (2004). When Cats Reigned Like Kings: On the Trail of the Sacred Cats. Kansas City, Missouri: Andrews McMeel. ISBN 0-7407-4697-9.
Jump up ^ Minou Reeves (2000). Muhammad in Europe. New York University (NYU) Press. p. 52. ISBN 0-8147-7533-0.
Jump up ^ Al-Thahabi, Shamsuddin. "Biography of al-Rifai". سير أعلام النبلاء (in Arabic). Archived from the original on 25 October 2014. Retrieved 11 November 2014.
Jump up ^ "Abu Hurairah and Cats". 13 January 2015.
Jump up ^ "Are Black Cats Really Bad Luck? [Hoax]". SocialNewsDaily. Archived from the original on 22 December 2015. Retrieved 19 December 2015.
Jump up ^ Davies, Norman (1996). Europe: A History. Oxford University Press. p. 543. ISBN 0-198-20171-0.
Jump up ^ Frazer, Sir James George. The Golden Bough, (1922). Online version. Archived 8 December 2006 at the Wayback Machine.
Jump up ^ Nora Sugobono (7 March 2010). "Las vidas del gato" (in Spanish). El Comercio. Archived from the original on 27 January 2012. Retrieved 19 March 2010.
Jump up ^ "Qual é a origem da lenda de que os gatos teriam sete vidas?". Mundo Estranho. Editora Abril. Archived from the original on 17 November 2015. Retrieved 15 November 2015.
Jump up ^ Tim Dowling (19 March 2010). "Tall tails: Pet myths busted". The Guardian. UK. Archived from the original on 9 September 2013. Retrieved 18 March 2010.
Jump up ^ ASPCA (30 June 2005). "The ASPCA Warns About High-Rise Falls by Cats". New York: About.com. Archived from the original on 22 May 2012. Retrieved 26 April 2012.
External links
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"Cat, Domestic, The". Encyclopedia Americana. 1920.
High-Resolution Images of the Cat Brain
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[hide]
v
t
e
Domestic cats
Felinology
Anatomy
Genetics
Dwarf cat
Kitten
Odd-eyed cat
Squitten
Coat genetics
Bicolor cat
Black cat
Calico cat
Tabby cat
Tortoiseshell cat
Health
Aging
Declawing
Diet
Neutering
Spaying
Vaccination
Behavior
Body language
Catfight
Catnip
Communication (Meow)
Kneading
Intelligence
Play and toys
Purr
Righting reflex
Senses
Human–cat
interaction
Ailurophobia
Animal-assisted therapy
Cat cafés
Cat massage
Cat meat
Cat-scratch disease
Cat show
Cats in ancient Egypt
Cultural depictions
Farm cat
Feral cat
Cats and Islam
Puppy cat
Ship's cat
Zoonosis
Registries
American Cat Fanciers Association
Associazione Nazionale Felina Italiana
Canadian Cat Association
Cat Aficionado Association
Cat Fanciers' Association
Fédération Internationale Féline
Governing Council of the Cat Fancy
Southern Africa Cat Council
The International Cat Association
World Cat Congress
World Cat Federation
Breeds
(full list)
(experimental breeds)
Fully domestic
Abyssinian
American Curl
American Shorthair
Balinese
Brazilian Shorthair
British Shorthair
Birman
Bombay
Burmese
California Spangled
Chartreux
Chinese Li Hua
Colorpoint Shorthair
Cornish Rex
Cymric
Devon Rex
Donskoy
Egyptian Mau
European Shorthair
Exotic Shorthair
German Rex
Himalayan
Japanese Bobtail
Javanese
Khao Manee
Korat
Kurilian Bobtail
Maine Coon
Manx
Munchkin
Norwegian Forest
Ocicat
Oriental Shorthair
Persian
Peterbald
Pixie-bob
Raas
Ragdoll
Ragamuffin
Russian Blue
Scottish Fold
Selkirk Rex
Siamese
Siberian
Singapura
Snowshoe
Somali
Sphynx
Thai
Traditional Persian
Tonkinese
Toyger
Turkish Angora
Turkish Van
Hybrid
Bengal
Chausie
Highlander
Savannah
Serengeti
Landraces
Aegean
Cyprus
Domestic long-haired
Domestic short-haired
Kellas
Sokoke
Van
Diseases and
disorders
Acne
Asthma
Calicivirus
Congenital sensorineural deafness
Flea
Heartworm
Hepatic lipidosis
Hypertrophic cardiomyopathy
Immunodeficiency virus
Infectious peritonitis
Leukemia virus
Lower urinary tract disease
Panleukopenia
Polydactyly
Rabies
Ringworm
Roundworm
Skin disorders
Tick
Toxoplasmosis
Viral rhinotracheitis
Book
Category
Portal
[show]
v
t
e
Extant Carnivora species
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Infraclass: Eutheria
Superorder: Laurasiatheria
[show]
Suborder Feliformia
Nandiniidae
Nandinia
African palm civet (N. binotata)
Herpestidae
(Mongooses)
Atilax
Marsh mongoose (A. paludinosus)
Bdeogale
Bushy-tailed mongoose (B. crassicauda)
Jackson's mongoose (B. jacksoni)
Black-footed mongoose (B. nigripes)
Crossarchus
Alexander's kusimanse (C. alexandri)
Angolan kusimanse (C. ansorgei)
Common kusimanse (C. obscurus)
Flat-headed kusimanse (C. platycephalus)
Cynictis
Yellow mongoose (C. penicillata)
Dologale
Pousargues's mongoose (D. dybowskii)
Galerella
Angolan slender mongoose (G. flavescens)
Black mongoose (G. nigrata)
Somalian slender mongoose (G. ochracea)
Cape gray mongoose (G. pulverulenta)
Slender mongoose (G. sanguinea)
Helogale
Ethiopian dwarf mongoose (H. hirtula)
Common dwarf mongoose (H. parvula)
Herpestes
Short-tailed mongoose (H. brachyurus)
Indian gray mongoose (H. edwardsii)
Indian brown mongoose (H. fuscus)
Egyptian mongoose (H. ichneumon)
Small Asian mongoose (H. javanicus)
Long-nosed mongoose (H. naso)
Collared mongoose (H. semitorquatus)
Ruddy mongoose (H. smithii)
Crab-eating mongoose (H. urva)
Stripe-necked mongoose (H. vitticollis)
Ichneumia
White-tailed mongoose (I. albicauda)
Liberiictus
Liberian mongoose (L. kuhni)
Mungos
Gambian mongoose (M. gambianus)
Banded mongoose (M. mungo)
Paracynictis
Selous' mongoose (P. selousi)
Rhynchogale
Meller's mongoose (R. melleri)
Suricata
Meerkat (S. suricatta)
Hyaenidae
(Hyenas)
Crocuta
Spotted hyena (C. crocuta)
Hyaena
Brown hyena (H. brunnea)
Striped hyena (H. hyaena)
Proteles
Aardwolf (P. cristatus)
Felidae
Large family listed below
Viverridae
Large family listed below
Eupleridae
Small family listed below
[show]
Family Felidae
Felinae
Acinonyx
Cheetah (A. jubatus)
Caracal
Caracal (C. caracal)
African golden cat (C. aurata)
Catopuma
Bay cat (C. badia)
Asian golden cat (C. temminckii)
Felis
European wildcat (F. silvestris)
African wildcat (F. lybica)
Jungle cat (F. chaus)
Black-footed cat (F. nigripes)
Sand cat (F. margarita)
Chinese mountain cat (F. bieti)
Domestic cat (F. catus)
Leopardus
Ocelot (L. pardalis)
Margay (L. wiedii)
Pampas cat (L. colocola)
Geoffroy's cat (L. geoffroyi)
Kodkod (L. guigna)
Andean mountain cat (L. jacobita)
Oncilla (L. tigrinus)
Southern tigrina (L. guttulus)
Leptailurus
Serval (L. serval)
Lynx
Canadian lynx (L. canadensis)
Eurasian lynx (L. lynx)
Iberian lynx (L. pardinus)
Bobcat (L. rufus)
Otocolobus
Pallas's cat (O. manul)
Pardofelis
Marbled cat (P. marmorata)
Prionailurus
Fishing cat (P. viverrinus)
Leopard cat (P. bengalensis)
Sundaland leopard cat (P. javanensis)
Flat-headed cat (P. planiceps)
Rusty-spotted cat (P. rubiginosus)
Puma
Cougar (P. concolor)
Herpailurus
Jaguarundi (H. yagouaroundi)
Pantherinae
Panthera
Lion (P. leo)
Jaguar (P. onca)
Leopard (P. pardus)
Tiger (P. tigris)
Snow leopard (P. uncia)
Neofelis
Clouded leopard (N. nebulosa)
Sunda clouded leopard (N. diardi)
[show]
Family Viverridae (includes Civets)
Paradoxurinae
Arctictis
Binturong (A. binturong)
Arctogalidia
Small-toothed palm civet (A. trivirgata)
Macrogalidia
Sulawesi palm civet (M. musschenbroekii)
Paguma
Masked palm civet (P. larvata)
Paradoxurus
Golden wet-zone palm civet (P. aureus)
Asian palm civet (P. hermaphroditus)
Jerdon's palm civet (P. jerdoni)
Golden palm civet (P. zeylonensis)
Hemigalinae
Chrotogale
Owston's palm civet (C. owstoni)
Cynogale
Otter civet (C. bennettii)
Diplogale
Hose's palm civet (D. hosei)
Hemigalus
Banded palm civet (H. derbyanus)
Prionodontinae
(Asiatic linsangs)
Prionodon
Banded linsang (P. linsang)
Spotted linsang (P. pardicolor)
Viverrinae
Civettictis
African civet (C. civetta)
Genetta
(Genets)
Abyssinian genet (G. abyssinica)
Angolan genet (G. angolensis)
Bourlon's genet (G. bourloni)
Crested servaline genet (G. cristata)
Common genet (G. genetta)
Johnston's genet (G. johnstoni)
Rusty-spotted genet (G. maculata)
Pardine genet (G. pardina)
Aquatic genet (G. piscivora)
King genet (G. poensis)
Servaline genet (G. servalina)
Haussa genet (G. thierryi)
Cape genet (G. tigrina)
Giant forest genet (G. victoriae)
Poiana
African linsang (P. richardsonii)
Leighton's linsang (P. leightoni)
Viverra
Malabar large-spotted civet (V. civettina)
Large-spotted civet (V. megaspila)
Malayan civet (V. tangalunga)
Large Indian civet (V. zibetha)
Viverricula
Small Indian civet (V. indica)
[show]
Family Eupleridae
Euplerinae
Cryptoprocta
Fossa (C. ferox)
Eupleres
Eastern falanouc (E. goudotii)
Western falanouc (E. major)
Fossa
Malagasy civet (F. fossana)
Galidiinae
Galidia
Ring-tailed mongoose (G. elegans)
Galidictis
Broad-striped Malagasy mongoose (G. fasciata)
Grandidier's mongoose (G. grandidieri)
Mungotictis
Narrow-striped mongoose (M. decemlineata)
Salanoia
Brown-tailed mongoose (S. concolor)
Durrell's vontsira (S. durrelli)
[show]
Suborder Caniformia (cont. below)
Ursidae
(Bears)
Ailuropoda
Giant panda (A. melanoleuca)
Helarctos
Sun bear (H. malayanus)
Melursus
Sloth bear (M. ursinus)
Tremarctos
Spectacled bear (T. ornatus)
Ursus
American black bear (U. americanus)
Brown bear (U. arctos)
Polar bear (U. maritimus)
Asian black bear (U. thibetanus)
Mephitidae
Conepatus
(Hog-nosed
skunks)
Molina's hog-nosed skunk (C. chinga)
Humboldt's hog-nosed skunk (C. humboldtii)
American hog-nosed skunk (C. leuconotus)
Striped hog-nosed skunk (C. semistriatus)
Mephitis
Hooded skunk (M. macroura)
Striped skunk (M. mephitis)
Mydaus
Sunda stink badger (M. javanensis)
Palawan stink badger (M. marchei)
Spilogale
(Spotted skunks)
Southern spotted skunk (S. angustifrons)
Western spotted skunk (S. gracilis)
Eastern spotted skunk (S. putorius)
Pygmy spotted skunk (S. pygmaea)
Procyonidae
Bassaricyon
(Olingos)
Eastern lowland olingo (B. alleni)
Northern olingo (B. gabbii)
Western lowland olingo (B. medius)
Olinguito (B. neblina)
Bassariscus
Ring-tailed cat (B. astutus)
Cacomistle (B. sumichrasti)
Nasua
(Coatis inclusive)
White-nosed coati (N. narica)
South American coati (N. nasua)
Nasuella
(Coatis inclusive)
Western mountain coati (N. olivacea)
Eastern mountain coati (N. meridensis)
Potos
Kinkajou (P. flavus)
Procyon
Crab-eating raccoon (P. cancrivorus)
Raccoon (P. lotor)
Cozumel raccoon (P. pygmaeus)
Ailuridae
Ailurus
Red panda (A. fulgens)
[show]
Suborder Caniformia (cont. above)
Otariidae
(Eared seals)
(includes fur seals
and sea lions)
(Pinniped inclusive)
Arctocephalus
South American fur seal (A. australis)
Australasian fur seal (A. forsteri)
Galápagos fur seal (A. galapagoensis)
Antarctic fur seal (A. gazella)
Juan Fernández fur seal (A. philippii)
Brown fur seal (A. pusillus)
Guadalupe fur seal (A. townsendi)
Subantarctic fur seal (A. tropicalis)
Callorhinus
Northern fur seal (C. ursinus)
Eumetopias
Steller sea lion (E. jubatus)
Neophoca
Australian sea lion (N. cinerea)
Otaria
South American sea lion (O. flavescens)
Phocarctos
New Zealand sea lion (P. hookeri)
Zalophus
California sea lion (Z. californianus)
Galápagos sea lion (Z. wollebaeki)
Odobenidae
(Pinniped inclusive)
Odobenus
Walrus (O. rosmarus)
Phocidae
(Earless seals)
(Pinniped inclusive)
Cystophora
Hooded seal (C. cristata)
Erignathus
Bearded seal (E. barbatus)
Halichoerus
Gray seal (H. grypus)
Histriophoca
Ribbon seal (H. fasciata)
Hydrurga
Leopard seal (H. leptonyx)
Leptonychotes
Weddell seal (L. weddellii)
Lobodon
Crabeater seal (L. carcinophagus)
Mirounga
(Elephant seals)
Northern elephant seal (M. angustirostris)
Southern elephant seal (M. leonina)
Monachus
Mediterranean monk seal (M. monachus)
Hawaiian monk seal (M. schauinslandi)
Ommatophoca
Ross seal (O. rossi)
Pagophilus
Harp seal (P. groenlandicus)
Phoca
Spotted seal (P. largha)
Harbor seal (P. vitulina)
Pusa
Caspian seal (P. caspica)
Ringed seal (P. hispida)
Baikal seal (P. sibirica)
Canidae
Large family listed below
Mustelidae
Large family listed below
[show]
Family Canidae (includes dogs)
Atelocynus
Short-eared dog (A. microtis)
Canis
Side-striped jackal (C. adustus)
African golden wolf (C. anthus)
Golden jackal (C. aureus)
Coyote (C. latrans)
Gray wolf (C. lupus)
Black-backed jackal (C. mesomelas)
Red wolf (C. rufus)
Ethiopian wolf (C. simensis)
Cerdocyon
Crab-eating fox (C. thous)
Chrysocyon
Maned wolf (C. brachyurus)
Cuon
Dhole (C. alpinus)
Lycalopex
Culpeo (L. culpaeus)
Darwin's fox (L. fulvipes)
South American gray fox (L. griseus)
Pampas fox (L. gymnocercus)
Sechuran fox (L. sechurae)
Hoary fox (L. vetulus)
Lycaon
African wild dog (L. pictus)
Nyctereutes
Raccoon dog (N. procyonoides)
Otocyon
Bat-eared fox (O. megalotis)
Speothos
Bush dog (S. venaticus)
Urocyon
Gray fox (U. cinereoargenteus)
Island fox (U. littoralis)
Vulpes
(Foxes)
Bengal fox (V. bengalensis)
Blanford's fox (V. cana)
Cape fox (V. chama)
Corsac fox (V. corsac)
Tibetan sand fox (V. ferrilata)
Arctic fox (V. lagopus)
Kit fox (V. macrotis)
Pale fox (V. pallida)
Rüppell's fox (V. rueppelli)
Swift fox (V. velox)
Red fox (V. vulpes)
Fennec fox (V. zerda)
[show]
Family Mustelidae
Lutrinae
(Otters)
Aonyx
African clawless otter (A. capensis)
Oriental small-clawed otter (A. cinerea)
Enhydra
Sea otter (E. lutris)
Hydrictis
Spotted-necked otter (H. maculicollis)
Lontra
North American river otter (L. canadensis)
Marine otter (L. felina)
Neotropical otter (L. longicaudis)
Southern river otter (L. provocax)
Lutra
Eurasian otter (L. lutra)
Hairy-nosed otter (L. sumatrana)
Lutrogale
Smooth-coated otter (L. perspicillata)
Pteronura
Giant otter (P. brasiliensis)
Mustelinae
(including badgers)
Arctonyx
Hog badger (A. collaris)
Eira
Tayra (E. barbara)
Galictis
Lesser grison (G. cuja)
Greater grison (G. vittata)
Gulo
Wolverine (G. gulo)
Ictonyx
Saharan striped polecat (I. libyca)
Striped polecat (I. striatus)
Lyncodon
Patagonian weasel (L. patagonicus)
Martes
(Martens)
American marten (M. americana)
Yellow-throated marten (M. flavigula)
Beech marten (M. foina)
Nilgiri marten (M. gwatkinsii)
European pine marten (M. martes)
Japanese marten (M. melampus)
Sable (M. zibellina)
Pekania
Fisher (P. pennanti)
Meles
Japanese badger (M. anakuma)
Asian badger (M. leucurus)
European badger (M. meles)
Mellivora
Honey badger (M. capensis)
Melogale
(Ferret-badgers)
Bornean ferret-badger (M. everetti)
Chinese ferret-badger (M. moschata)
Javan ferret-badger (M. orientalis)
Burmese ferret-badger (M. personata)
Mustela
(Weasels and Ferrets)
Amazon weasel (M. africana)
Mountain weasel (M. altaica)
Stoat (M. erminea)
Steppe polecat (M. eversmannii)
Colombian weasel (M. felipei)
Long-tailed weasel (M. frenata)
Japanese weasel (M. itatsi)
Yellow-bellied weasel (M. kathiah)
European mink (M. lutreola)
Indonesian mountain weasel (M. lutreolina)
Black-footed ferret (M. nigripes)
Least weasel (M. nivalis)
Malayan weasel (M. nudipes)
European polecat (M. putorius)
Siberian weasel (M. sibirica)
Back-striped weasel (M. strigidorsa)
Egyptian weasel (M. subpalmata)
Neovison
(Minks)
American mink (N. vison)
Poecilogale
African striped weasel (P. albinucha)
Taxidea
American badger (T. taxus)
Vormela
Marbled polecat (V. peregusna)
Taxon identifiers
Wd: Q20980826
EoL: 4443860
EPPO: FELIDO
Fossilworks: 104159
GBIF: 2435035
iNaturalist: 118552
IRMNG: 10196305
ITIS: 183798
MSW: 14000031
NCBI: 9685
NZOR: 7d7d7c68-baa8-4908-bdc4-b747950f6318
Authority control
GND: 4030046-8
HDS: 41559
NARA: 10647397
Retrieved from "https://en.wikipedia.org/w/index.php?title=Cat&oldid=842123260"
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Domesticated animals
Cats
Mammals described in 1758
Cosmopolitan vertebrates
Invasive mammal species
Vertebrate animal models
Cat diseases
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interlanguage-link-target

values:
interlanguage-link interwiki-zu
display: none;

values:
interlanguage-link interwiki-is
display: none;

values:
interlanguage-link interwiki-it
display: none;

values: חתול הבית – Hebrew

values:
interlanguage-link interwiki-he
display: none;

values:
https://jv.wikipedia.org/wiki/Kucing
Kucing – Javanese
jv
jv
interlanguage-link-target

values:
interlanguage-link interwiki-jv
display: none;

values: ಬೆಕ್ಕು – Kannada

values:
interlanguage-link interwiki-kn
display: none;

values:
https://pam.wikipedia.org/wiki/Pusa
Pusa – Pampanga
pam
pam
interlanguage-link-target

values:
interlanguage-link interwiki-pam
display: none;

values: შინაური კატა – Georgian

values:
interlanguage-link interwiki-ka
display: none;

values:
interlanguage-link interwiki-ks
display: none;

values:
https://csb.wikipedia.org/wiki/K%C3%B2t
Kòt – Kashubian
csb
csb
interlanguage-link-target

values:
interlanguage-link interwiki-csb
display: none;

values: Мысық – Kazakh

values:
interlanguage-link interwiki-kk
display: none;

values:
https://rw.wikipedia.org/wiki/Injangwe
Injangwe – Kinyarwanda
rw
rw
interlanguage-link-target

values:
interlanguage-link interwiki-rw
display: none;

values:
https://sw.wikipedia.org/wiki/Paka-kaya
Paka-kaya – Swahili
sw
sw
interlanguage-link-target

values:
interlanguage-link interwiki-sw
display: none;

values:
interlanguage-link interwiki-kv
display: none;

values:
https://kg.wikipedia.org/wiki/Nyawu
Nyawu – Kongo
kg
kg
interlanguage-link-target

values:
interlanguage-link interwiki-kg
display: none;

values:
https://ht.wikipedia.org/wiki/Chat
Chat – Haitian Creole
ht
ht
interlanguage-link-target

values:
interlanguage-link interwiki-ht
display: none;

values:
interlanguage-link interwiki-ku
display: none;

values: Бакма мышыктар – Kyrgyz

values:
interlanguage-link interwiki-ky
display: none;

values:
interlanguage-link interwiki-lbe
display: none;

values:
interlanguage-link interwiki-lez
display: none;

values:
interlanguage-link interwiki-lo
display: none;

values:
interlanguage-link interwiki-ltg
display: none;

values:
https://la.wikipedia.org/wiki/Feles
Feles – Latin
la
la
interlanguage-link-target

values: interlanguage-link interwiki-la badge-Q17437796 badge-featuredarticle

values:
interlanguage-link interwiki-lv
display: none;

values:
https://lb.wikipedia.org/wiki/Hauskaz
Hauskaz – Luxembourgish
lb
lb
interlanguage-link-target

values:
interlanguage-link interwiki-lb
display: none;

values:
https://lt.wikipedia.org/wiki/Kat%C4%97
Katė – Lithuanian
lt
lt
interlanguage-link-target

values: interlanguage-link interwiki-lt badge-Q17437798 badge-goodarticle

values:
interlanguage-link interwiki-lij
display: none;

values:
https://li.wikipedia.org/wiki/Hoeskat
Hoeskat – Limburgish
li
li
interlanguage-link-target

values:
interlanguage-link interwiki-li
display: none;

values:
interlanguage-link interwiki-ln
display: none;

values:
interlanguage-link interwiki-olo
display: none;

values:
https://jbo.wikipedia.org/wiki/mlatu
mlatu – Lojban
jbo
jbo
interlanguage-link-target

values:
interlanguage-link interwiki-jbo
display: none;

values:
interlanguage-link interwiki-lmo
display: none;

values:
https://hu.wikipedia.org/wiki/Macska
Macska – Hungarian
hu
hu
interlanguage-link-target

values: interlanguage-link interwiki-hu badge-Q17437796 badge-featuredarticle

values:
interlanguage-link interwiki-mk
display: none;

values:
https://mg.wikipedia.org/wiki/Saka
Saka – Malagasy
mg
mg
interlanguage-link-target

values:
interlanguage-link interwiki-mg
display: none;

values: interlanguage-link interwiki-ml badge-Q17437796 badge-featuredarticle

values:
https://mt.wikipedia.org/wiki/Qattus
Qattus – Maltese
mt
mt
interlanguage-link-target

values:
interlanguage-link interwiki-mt
display: none;

values:
interlanguage-link interwiki-mr
display: none;

values:
interlanguage-link interwiki-arz
display: none;

values:
interlanguage-link interwiki-mzn
display: none;

values:
https://ms.wikipedia.org/wiki/Kucing
Kucing – Malay
ms
ms
interlanguage-link-target

values: interlanguage-link interwiki-ms badge-Q17437796 badge-featuredarticle

values:
https://cdo.wikipedia.org/wiki/M%C3%A0
Mà – Min Dong Chinese
cdo
cdo
interlanguage-link-target

values:
interlanguage-link interwiki-cdo
display: none;

values:
https://mwl.wikipedia.org/wiki/Gato
Gato – Mirandese
mwl
mwl
interlanguage-link-target

values:
interlanguage-link interwiki-mwl
display: none;

values:
interlanguage-link interwiki-mdf
display: none;

values:
interlanguage-link interwiki-mn
display: none;

values: ကြောင် (တိရစ္ဆာန်) – Burmese

values:
interlanguage-link interwiki-my
display: none;

values:
https://nah.wikipedia.org/wiki/Mizton
Mizton – Nāhuatl
nah
nah
interlanguage-link-target

values:
interlanguage-link interwiki-nah
display: none;

values:
https://na.wikipedia.org/wiki/Butsi
Butsi – Nauru
interlanguage-link-target

values:
interlanguage-link interwiki-na
display: none;

values:
interlanguage-link interwiki-nl
display: none;

values:
https://nds-nl.wikipedia.org/wiki/Kat
Kat – Low Saxon
nds-NL
nds-NL
interlanguage-link-target

values:
interlanguage-link interwiki-nds-nl
display: none;

values: बिरालो – Nepali

values:
interlanguage-link interwiki-ne
display: none;

values:
interlanguage-link interwiki-new
display: none;

values:
interlanguage-link interwiki-ja
display: none;

values:
interlanguage-link interwiki-ce
display: none;

values:
https://frr.wikipedia.org/wiki/Kaat
Kaat – Northern Frisian
frr
frr
interlanguage-link-target

values:
interlanguage-link interwiki-frr
display: none;

values:
https://no.wikipedia.org/wiki/Tamkatt
Tamkatt – Norwegian
no
no
interlanguage-link-target

values:
interlanguage-link interwiki-no
display: none;

values:
https://nn.wikipedia.org/wiki/Katt
Katt – Norwegian Nynorsk
nn
nn
interlanguage-link-target

values:
interlanguage-link interwiki-nn
display: none;

values:
https://nrm.wikipedia.org/wiki/Cat
Cat – Norman
nrm
nrm
interlanguage-link-target

values:
interlanguage-link interwiki-nrm
display: none;

values:
interlanguage-link interwiki-nov
display: none;

values:
interlanguage-link interwiki-oc
display: none;

values:
interlanguage-link interwiki-mhr
display: none;

values:
https://uz.wikipedia.org/wiki/Mushuk
Mushuk – Uzbek
uz
uz
interlanguage-link-target

values:
interlanguage-link interwiki-uz
display: none;

values:
interlanguage-link interwiki-pa
display: none;

values:
interlanguage-link interwiki-pnb
display: none;

values:
interlanguage-link interwiki-ps
display: none;

values:
https://jam.wikipedia.org/wiki/Pus
Pus – Jamaican Creole English
jam
jam
interlanguage-link-target

values:
interlanguage-link interwiki-jam
display: none;

values:
interlanguage-link interwiki-koi
display: none;

values: ឆ្មា – Khmer

values:
interlanguage-link interwiki-km
display: none;

values:
https://pcd.wikipedia.org/wiki/Cot
Cot – Picard
pcd
pcd
interlanguage-link-target

values:
interlanguage-link interwiki-pcd
display: none;

values:
https://pms.wikipedia.org/wiki/Gat
Gat – Piedmontese
pms
pms
interlanguage-link-target

values:
interlanguage-link interwiki-pms
display: none;

values:
https://nds.wikipedia.org/wiki/Katt
Katt – Low German
nds
nds
interlanguage-link-target

values:
interlanguage-link interwiki-nds
display: none;

values:
interlanguage-link interwiki-pl
display: none;

values:
https://pt.wikipedia.org/wiki/Gato
Gato – Portuguese
pt
pt
interlanguage-link-target

values:
interlanguage-link interwiki-pt
display: none;

values:
interlanguage-link interwiki-ro
display: none;

values:
https://rmy.wikipedia.org/wiki/Magodja
Magodja – Romani
rmy
rmy
interlanguage-link-target

values:
interlanguage-link interwiki-rmy
display: none;

values:
https://rm.wikipedia.org/wiki/Giat
Giat – Romansh
rm
rm
interlanguage-link-target

values:
interlanguage-link interwiki-rm
display: none;

values:
https://qu.wikipedia.org/wiki/Michi
Michi – Quechua
qu
qu
interlanguage-link-target

values:
interlanguage-link interwiki-qu
display: none;

values: Мачка – Rusyn

values:
interlanguage-link interwiki-rue
display: none;

values: interlanguage-link interwiki-ru badge-Q17437796 badge-featuredarticle

values: Куоска – Sakha

values:
interlanguage-link interwiki-sah
display: none;

values: मार्जालः – Sanskrit

values:
interlanguage-link interwiki-sa
display: none;

values:
https://sc.wikipedia.org/wiki/Felis
Felis – Sardinian
sc
sc
interlanguage-link-target

values:
interlanguage-link interwiki-sc
display: none;

values:
https://sco.wikipedia.org/wiki/Cat
Cat – Scots
sco
sco
interlanguage-link-target

values:
interlanguage-link interwiki-sco
display: none;

values:
https://stq.wikipedia.org/wiki/Kat
Kat – Saterland Frisian
stq
stq
interlanguage-link-target

values:
interlanguage-link interwiki-stq
display: none;

values:
https://st.wikipedia.org/wiki/Katse
Katse – Southern Sotho
st
st
interlanguage-link-target

values:
interlanguage-link interwiki-st
display: none;

values:
https://nso.wikipedia.org/wiki/Katse
Katse – Northern Sotho
nso
nso
interlanguage-link-target

values:
interlanguage-link interwiki-nso
display: none;

values:
https://sq.wikipedia.org/wiki/Macja
Macja – Albanian
sq
sq
interlanguage-link-target

values:
interlanguage-link interwiki-sq
display: none;

values:
https://scn.wikipedia.org/wiki/Jattu
Jattu – Sicilian
scn
scn
interlanguage-link-target

values:
interlanguage-link interwiki-scn
display: none;

values:
interlanguage-link interwiki-si
display: none;

values:
https://simple.wikipedia.org/wiki/Cat
Cat – Simple English
simple
simple
interlanguage-link-target

values:
interlanguage-link interwiki-simple
display: none;

values:
interlanguage-link interwiki-sd
display: none;

values:
https://ss.wikipedia.org/wiki/Likati
Likati – Swati
ss
ss
interlanguage-link-target

values:
interlanguage-link interwiki-ss
display: none;

values: interlanguage-link interwiki-sk badge-Q17437796 badge-featuredarticle

values:
interlanguage-link interwiki-sl
display: none;

values:
interlanguage-link interwiki-cu
display: none;

values:
https://szl.wikipedia.org/wiki/Kot
Kot – Silesian
szl
szl
interlanguage-link-target

values:
interlanguage-link interwiki-szl
display: none;

values:
https://so.wikipedia.org/wiki/Yaanyuur
Yaanyuur – Somali
so
so
interlanguage-link-target

values:
interlanguage-link interwiki-so
display: none;

values:
interlanguage-link interwiki-ckb
display: none;

values:
https://srn.wikipedia.org/wiki/Puspusi
Puspusi – Sranan Tongo
srn
srn
interlanguage-link-target

values:
interlanguage-link interwiki-srn
display: none;

values: interlanguage-link interwiki-sr badge-Q17437796 badge-featuredarticle

values:
interlanguage-link interwiki-sh
display: none;

values:
https://su.wikipedia.org/wiki/Ucing
Ucing – Sundanese
su
su
interlanguage-link-target

values:
interlanguage-link interwiki-su
display: none;

values:
https://fi.wikipedia.org/wiki/Kissa
Kissa – Finnish
fi
fi
interlanguage-link-target

values:
interlanguage-link interwiki-fi
display: none;

values:
https://sv.wikipedia.org/wiki/Katt
Katt – Swedish
sv
sv
interlanguage-link-target

values:
interlanguage-link interwiki-sv
display: none;

values:
https://tl.wikipedia.org/wiki/Pusa
Pusa – Tagalog
tl
tl
interlanguage-link-target

values:
interlanguage-link interwiki-tl
display: none;

values: பூனை – Tamil

values:
interlanguage-link interwiki-ta
display: none;

values:
https://kab.wikipedia.org/wiki/Amcic
Amcic – Kabyle
kab
kab
interlanguage-link-target

values:
interlanguage-link interwiki-kab
display: none;

values: Мәче – Tatar

values:
interlanguage-link interwiki-tt
display: none;

values: పిల్లి – Telugu

values:
interlanguage-link interwiki-te
display: none;

values:
interlanguage-link interwiki-th
display: none;

values: Гурба – Tajik

values:
interlanguage-link interwiki-tg
display: none;

values:
https://to.wikipedia.org/wiki/Pusi
Pusi – Tongan
to
to
interlanguage-link-target

values:
interlanguage-link interwiki-to
display: none;

values:
interlanguage-link interwiki-chr
display: none;

values: ಪುಚ್ಚೆ – Tulu

values:
interlanguage-link interwiki-tcy
display: none;

values:
https://tr.wikipedia.org/wiki/Kedi
Kedi – Turkish
tr
tr
interlanguage-link-target

values:
interlanguage-link interwiki-tr
display: none;

values: Коӵыш – Udmurt

values:
interlanguage-link interwiki-udm
display: none;

values:
https://bug.wikipedia.org/wiki/Meong
Meong – Buginese
bug
bug
interlanguage-link-target

values:
interlanguage-link interwiki-bug
display: none;

values: Кіт свійський – Ukrainian

values:
interlanguage-link interwiki-uk
display: none;

values:
interlanguage-link interwiki-ur
display: none;

values: مۈشۈك – Uyghur

values:
interlanguage-link interwiki-ug
display: none;

values:
https://za.wikipedia.org/wiki/Meuz
Meuz – Zhuang
za
za
interlanguage-link-target

values:
interlanguage-link interwiki-za
display: none;

values:
https://vec.wikipedia.org/wiki/Gato
Gato – Venetian
vec
vec
interlanguage-link-target

values:
interlanguage-link interwiki-vec
display: none;

values:
interlanguage-link interwiki-vep
display: none;

values:
https://vi.wikipedia.org/wiki/M%C3%A8o
Mèo – Vietnamese
vi
vi
interlanguage-link-target

values:
interlanguage-link interwiki-vi
display: none;

values:
https://fiu-vro.wikipedia.org/wiki/Kass
Kass – Võro
fiu-vro
fiu-vro
interlanguage-link-target

values:
interlanguage-link interwiki-fiu-vro
display: none;

values:
https://wa.wikipedia.org/wiki/Tchet
Tchet – Walloon
wa
wa
interlanguage-link-target

values:
interlanguage-link interwiki-wa
display: none;

values:
https://vls.wikipedia.org/wiki/Katte
Katte – West Flemish
vls
vls
interlanguage-link-target

values:
interlanguage-link interwiki-vls
display: none;

values:
https://war.wikipedia.org/wiki/Misay
Misay – Waray
war
war
interlanguage-link-target

values:
interlanguage-link interwiki-war
display: none;

values:
https://wo.wikipedia.org/wiki/Muus
Muus – Wolof
wo
wo
interlanguage-link-target

values:
interlanguage-link interwiki-wo
display: none;

values:
interlanguage-link interwiki-wuu
display: none;

values:
interlanguage-link interwiki-yi
display: none;

values:
interlanguage-link interwiki-yo
display: none;

values:
interlanguage-link interwiki-zh-yue
display: none;

values:
interlanguage-link interwiki-diq
display: none;

values:
interlanguage-link interwiki-bat-smg
display: none;

values:
https://zh.wikipedia.org/wiki/%E7%8C%AB
猫 – Chinese
zh
zh
interlanguage-link-target

values:
interlanguage-link interwiki-zh
display: none;

values:
https://gor.wikipedia.org/wiki/Tete
Tete – Gorontalo
gor
gor
interlanguage-link-target

values:
interlanguage-link interwiki-gor
display: none;

values:
interlanguage-link interwiki-inh
display: none;

values:
interlanguage-link interwiki-kbp
display: none;

values:
https://lfn.wikipedia.org/wiki/Gato
Gato – Lingua Franca Nova
lfn
lfn
interlanguage-link-target

values:
interlanguage-link interwiki-lfn
display: none;

values: All languages (initial selection from common choices by you and others)

values: mediawiki ltr sitedir-ltr mw-hide-empty-elt ns-0 ns-subject page-Cat rootpage-Cat skin-vector action-view