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History of Latin
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One of the seven ceiling frescoes painted by Bartolomeo Altomonte in his 80th year for the library of Admont Abbey. An allegory of the Enlightenment, it shows Aurora, goddess of dawn, with the geniuses of language in her train awakening Morpheus, god of dreaming, a symbol of man. The geniuses are Grammar, Didactic, Greek, Hebrew and Latin.
Approximate distribution of languages in Iron Age Italy during the 6th century BC. Latin is confined to Latium, a small region on the coast of west central Italy, hemmed in by other Italic peoples on the east and south and the powerful Etruscan civilization on the north.
Latin is a member of the broad family of Italic languages. Its alphabet, the Latin alphabet, emerged from the Old Italic alphabets, which in turn were derived from the Greek and Phoenician scripts. Historical Latin came from the prehistoric language of the Latium region, specifically around the River Tiber, where Roman civilization first developed. How and when Latin came to be spoken by the Romans are questions that have long been debated. Various influences on Latin of Celtic dialects in northern Italy, the non-Indo-European Etruscan language in Central Italy, and the Greek of southern Italy have been detected, but when these influences entered the native Latin is not known for certain.
Surviving Latin literature consists almost entirely of Classical Latin in its broadest definition. It includes a polished and sometimes highly stylized literary language sometimes termed Golden Latin, which spans the 1st century BC and the early years of the 1st century AD. However, throughout the history of ancient Rome the spoken language differed in both grammar and vocabulary from that of literature, and is referred to as Vulgar Latin. In addition to Latin, Greek was often spoken by the well-educated elite, who studied it in school and acquired Greek tutors from among the influx of enslaved educated Greek prisoners of war. In the eastern half of the Roman Empire, which became the Byzantine Empire, the Greek Koine of Hellenism remained current and was never replaced by Latin.
Contents
[hide]
1 Origins
2 Old Latin
3 Classical Latin
3.1 Golden Age
3.2 Silver Age
4 Late Latin
5 Vulgar Latin
6 Romance languages
7 Ecclesiastical Latin
8 Medieval Latin
9 Renaissance Latin
10 New Latin
11 Phonology
11.1 Vowels
11.1.1 Pure vowels
11.1.1.1 Initial syllables
11.1.1.2 Medial syllables
11.1.1.3 Syncope
11.1.1.4 Final syllables
11.1.2 Diphthongs
11.1.2.1 Initial syllables
11.1.2.2 Medial syllables
11.1.2.3 Final syllables
11.1.3 Syllabic resonants and laryngeals
11.2 Consonants
11.2.1 Aspirates
11.2.2 Labiovelars
11.2.3 Other sequences
11.2.4 S-rhotacism
12 See also
13 Notes
14 Sources
15 External links
Origins[edit]
Main article: Italic languages
The name Latin derives from the Italic tribal group named Latini that settled around the 10th century BC in Latium, and the dialect spoken by these people.[1]
The Italic languages form a centum subfamily of the Indo-European language family. These include the Romance, Germanic, Celtic, and Hellenic languages, and a number of extinct ones.
Broadly speaking, in initial syllables the Indo-European simple vowels — (*a), *e, *i, *o, *u; short and long — are usually retained in Latin. The schwa indogermanicum (*ə) appears in Latin as a (cf. IE *pəter > L pater). Diphthongs are also preserved in Old Latin, but in Classical Latin some tend to become monophthongs (for example oi > ū or oe, and ei > ē > ī).[2] In non-initial syllables, there was more vowel reduction. The most extreme case occurs with short vowels in medial open syllables (i.e. short vowels followed by at most a single consonant, occurring neither in the first nor last syllable): All are reduced to a single vowel, which appears as i in most cases, but e (sometimes o) before r, and u before an l which is followed by o or u. In final syllables, short e and o are usually raised to i and u, respectively.
Consonants are generally more stable. However, the Indo-European voiced aspirates bh, dh, gh, gwh are not maintained, becoming f, f, h, f respectively at the beginning of a word, but usually b, d, g, v elsewhere. Note that non-initial dh becomes b next to r or u, e.g. *h₁rudh- "red" > rub-, e.g. rubeō "to be red"; *werdh- "word" > verbum. s between vowels famously becomes r, e.g. flōs "flower", gen. flōris; erō "I will be" vs. root es-; aurōra "dawn" < *ausōsā (cf. Germanic *aust- > English "east", Vedic Sanskrit uṣā́s "dawn"); soror "sister" < *sosor < *swesōr (cf. Old English sweostor "sister").
Of the original eight cases of Proto-Indo-European, Latin inherited six: nominative, vocative, accusative, genitive, dative, and ablative. The Indo-European locative survived in the declensions of some place names and a few common nouns, such as Roma "Rome" (locative Romae) and domus "home" (locative domī "at home"). Vestiges of the instrumental case may remain in adverbial forms ending in -ē.[3]
It is believed that the earliest surviving inscription is a seventh-century B.C. pin known as the Praenestine fibula, which reads Manios med fhefhaked Numasioi "Manius made me for Numerius".[4]
Old Latin[edit]
The Duenos inscription, from the 6th century BC, is the second-earliest known Latin text.
Main article: Old Latin
Old Latin (also called Early Latin or Archaic Latin) refers to the period of Latin texts before the age of Classical Latin, extending from textual fragments that probably originated in the Roman monarchy to the written language of the late Roman republic about 75 BC. Almost all the writing of its earlier phases is inscriptional.
Some phonological characteristics of older Latin are the case endings -os and -om (later Latin -us and -um). In many locations, classical Latin turned intervocalic /s/ into /r/. This had implications for declension: early classical Latin, honos, honosis; Classical honor, honoris ("honor"). Some Latin texts preserve /s/ in this position, such as the Carmen Arvale's lases for lares.
Classical Latin[edit]
Main article: Classical Latin
Julius Caesar's Commentarii de Bello Gallico is one of the most famous classical Latin texts of the Golden Age of Latin. The unvarnished, journalistic style of this upper-class general has long been taught as a model of the urbane Latin officially spoken and written in the floruit of the Roman republic.
Classical Latin is the form of the Latin language used by the ancient Romans in Classical Latin literature. In the latest and narrowest philological model its use spanned the Golden Age of Latin literature – broadly the 1st century BC and the early 1st century AD – possibly extending to the Silver Age – broadly the 1st and 2nd centuries. It was a polished written literary language based on the refined spoken language of the upper classes. Classical Latin differs from Old Latin: the earliest inscriptional language and the earliest authors, such as Ennius, Plautus and others, in a number of ways; for example, the early -om and -os endings shifted into -um and -us ones, and some lexical differences also developed, such as the broadening of the meaning of words.[5] In the broadest and most ancient sense, the classical period includes the authors of Early Latin, the Golden Age and the Silver Age.
Golden Age[edit]
The golden age of Latin literature is a period consisting roughly of the time from 75 BC to AD 14, covering the end of the Roman Republic and the reign of Augustus Caesar. In the currently used philological model this period represents the peak of Latin literature. Since the earliest post-classical times the Latin of those authors has been an ideal norm of the best Latin, which other writers should follow.
Silver Age[edit]
In reference to Roman literature, the Silver age covers the first two centuries AD directly after the Golden age. Literature from the Silver Age is more embellished with mannerisms.
Late Latin[edit]
Main article: Late Latin
Late Latin is the administrative and literary language of Late Antiquity in the late Roman empire and states that succeeded the Western Roman Empire over the same range. By its broadest definition it is dated from about 200 AD to about 900 AD when it was replaced by written Romance languages. Opinion concerning whether it should be considered classical is divided. The authors of the period looked back to a classical period they believed should be imitated and yet their styles were often classical. According to the narrowest definitions, Late Latin did not exist and the authors of the times are to be considered medieval.
Vulgar Latin[edit]
Main article: Vulgar Latin
Vulgar Latin, as in this political graffito at Pompeii, was the language of the ordinary people of the Roman Empire, distinct from the Classical Latin of literature.
Vulgar Latin (in Latin, sermo vulgaris) is a blanket term covering vernacular dialects of the Latin language spoken from earliest times in Italy until the latest dialects of the Western Roman Empire, diverging still further, evolved into the early Romance languages – whose writings began to appear about the 9th century.
This spoken Latin differed from the literary language of Classical Latin in its grammar and vocabulary. It is likely to have evolved over time, with some features not appearing until the late Empire. Other features are likely to have been in place much earlier. Because there are few phonetic transcriptions of the daily speech of these Latin speakers (to match, for example, the post-classical Appendix Probi) Vulgar Latin must be studied mainly by indirect methods.
A replica of the Old Roman Cursive inspired by the Vindolanda tablets
Knowledge of Vulgar Latin comes from a variety of sources. First, the comparative method reconstructs items of the mother language from the attested Romance languages. Also, prescriptive grammar texts from the Late Latin period condemn some usages as errors, providing insight into how Latin was actually spoken. The solecisms and non-Classical usages occasionally found in late Latin texts also shed light on the spoken language. A windfall source lies in the chance finds of wax tablets such as those found at Vindolanda on Hadrian's Wall. The Roman cursive script was used on these tablets.
Romance languages[edit]
Main article: Romance languages
The Romance languages, a major branch of the Indo-European language family, comprise all languages that descended from Latin, the language of the Roman Empire. The Romance languages have more than 700 million native speakers worldwide, mainly in the Americas, Europe, and Africa, as well as in many smaller regions scattered through the world.
All Romance languages descend from Vulgar Latin, the language of soldiers, settlers, and slaves of the Roman Empire, which was substantially different from that of the Roman literati. Between 200 BC and AD 100, the expansion of the Empire and the administrative and educational policies of Rome made Vulgar Latin the dominant vernacular language over a wide area which stretched from the Iberian Peninsula to the west coast of the Black Sea. During the Empire's decline and after its collapse and fragmentation in the 5th century, Vulgar Latin began to evolve independently within each local area, and eventually diverged into dozens of distinct languages. The overseas empires established by Spain, Portugal and France after the 15th century then spread these languages to other continents – about two thirds of all Romance speakers are now outside Europe.
In spite of the multiple influences of pre-Roman languages and later invasions, the phonology, morphology, lexicon, and syntax of all Romance languages are predominantly derived from Vulgar Latin. As a result, the group shares a number of linguistic features that set it apart from other Indo-European branches.
Ecclesiastical Latin[edit]
Main article: Ecclesiastical Latin
Ecclesiastical Latin (sometimes called Church Latin) is a broad and analogous term referring to the Latin language as used in documents of the Roman Catholic Church, its liturgies (mainly in past times) and during some periods the preaching of its ministers. Ecclesiastical Latin is not a single style: the term merely means the language promulgated at any time by the church. In terms of stylistic periods, it belongs to Late Latin in the Late Latin period, Medieval Latin in the Medieval Period, and so on through to the present. One may say that, starting from the church's decision in the early Late Latin period to use a simple and unornamented language that would be comprehensible to ordinary Latin speakers and yet still be elegant and correct, church Latin is usually a discernible substyle within the major style of the period. Its authors in the New Latin period are typically paradigmatic of the best Latin and that is true in contemporary times. The decline in its use within the last 100 years has been a matter of regret to some, who have formed organizations inside and outside the church to support its use and to use it.
Medieval Latin[edit]
Main article: Medieval Latin
Page with medieval Latin text from the Carmina Cantabrigiensia (Cambridge University Library, Gg. 5. 35), 11. cent.
Medieval Latin refers to the literary and administrative Latin used in the Middle Ages. It exhibits much variation between individual authors, mainly due to poor communications in those times between different regions. The individuality is characterised by a different range of solecisms and by the borrowing of different words from Vulgar Latin or from the local vernacular. Some styles show features intermediate between Latin and Romance languages; others are closer to classical Latin. The stylistic variations came to an end with the rise of nations and new empires in the Renaissance period, and the authority of the first universities imposing a new style, Renaissance Latin.
Renaissance Latin[edit]
Main article: Renaissance Latin
Renaissance Latin is a name given to the Latin written during the European Renaissance in the 14th-16th centuries, particularly distinguished by the distinctive Latin style developed by the humanist movement.
Ad fontes was the general cry of the humanists, and as such their Latin style sought to purge Latin of the medieval Latin vocabulary and stylistic accretions that it had acquired in the centuries after the fall of the Roman Empire. They looked to Golden Age Latin literature, and especially to Cicero in prose and Virgil in poetry, as the arbiters of Latin style. They abandoned the use of the sequence and other accentual forms of meter, and sought instead to revive the Greek formats that were used in Latin poetry during the Roman period. The humanists condemned the large body of medieval Latin literature as "gothic" – for them, a term of abuse – and believed instead that only ancient Latin from the Roman period was "real Latin".
The humanists also sought to purge written Latin of medieval developments in its orthography. They insisted, for example, that ae be written out in full wherever it occurred in classical Latin; medieval scribes often wrote e instead of ae. They were much more zealous than medieval Latin writers in distinguishing t from c: because the effects of palatalization made them homophones, medieval scribes often wrote, for example, eciam for etiam. Their reforms even affected handwriting: humanists usually wrote Latin in a script derived from Carolingian minuscule, the ultimate ancestor of most contemporary lower-case typefaces, avoiding the black-letter scripts used in the Middle Ages. Erasmus even proposed that the then-traditional pronunciations of Latin be abolished in favour of his reconstructed version of classical Latin pronunciation.
The humanist plan to remake Latin was largely successful, at least in education. Schools now taught the humanistic spellings, and encouraged the study of the texts selected by the humanists, largely to the exclusion of later Latin literature. On the other hand, while humanist Latin was an elegant literary language, it became much harder to write books about law, medicine, science or contemporary politics in Latin while observing all of the humanists' norms of vocabulary purging and classical usage. Because humanist Latin lacked precise vocabulary to deal with modern issues, their reforms accelerated the transformation of Latin from a working language to an object of antiquarian study. Their attempts at literary work, especially poetry, often have a strong element of pastiche.
New Latin[edit]
A Recent Latin inscription at Salamanca University commemorating the visit of the then-Prince "Akihitus" and Princess "Michika" of Japan on 28 February 1985
Main article: New Latin
New Latin (or Neo-Latin) is a post-medieval version of Latin, now used primarily in international scientific vocabulary and systematics. The term came into widespread use towards the end of the 1890s among linguists and scientists.
Classicists use the term "Neo-Latin" to describe the use of the Latin language for any purpose, scientific or literary, after the Renaissance (for which purpose they often use the date 1500), although, for example, the editors of the I Tatti Renaissance Library call their Renaissance Latin language texts Neo-Latin as well. Such Contemporary Latin includes ecclesiastical use, as well as translations from modern languages into Latin and the occasional poetry. Under the name "Living Latin", some have advocated reviving the language as a means of spoken communication.
Phonology[edit]
Vowels[edit]
Proto-Italic inherited all ten of the early post-Proto-Indo-European simple vowels (i.e. at a time when laryngeals had colored and often lengthened adjacent vowels and then disappeared in many circumstances): *a, *e, *i, *o, *u, *ā, *ē, *ī, *ō, *ū. It also inherited all of the post-PIE diphthongs except for *eu, which became *ou. Proto-Italic and Old Latin had a stress accent on the first syllable of a word, and this caused steady reduction and eventual deletion of many short vowels in non-initial syllables while affecting initial syllables much less. Long vowels were largely unaffected in general except in final syllables, where they had a tendency to shorten.
Development of Proto-Italic vowels in Latin
Initial
Medial
Final
Proto-Italic
+r
+l pinguis
+other
+one consonant
+cluster
Absolutely final
one consonant
cluster
s
m, n
other
a
a e u i e i e i e e
e
e
i
i i
o
o u u
u
u e
ā
ā a a, ā
ē
ē e ē?
ī
ī i ī?
ō
ō o ō
ū
ū u ū?
ai
ae ī
au
au ū
ei
ī
oi
ū, oe ū ī
ou
ū
Note: For the following examples, it helps to keep in mind the normal correspondences between PIE and certain other languages:
Development of some Proto-Indo-European sounds in other languages
(post-)PIE
Ancient Greek
Old English
Gothic
Sanskrit
Notes
*i
i i i, aí /ɛ/ i
*e
e e i, aí /ɛ/ a
*a
a a a a
*o
o a a a
*u
u u, o u, aú /ɔ/ u
*ī
ī ī ei /ī/ ī
*ē
ē ā ē ā
*ā
ē (Attic);
ā (Doric, etc.) ō ō ā
*ō
ō ō ō ā
*ū
ū ū ū ū
*ai
ai ā ái ē
*au
au ēa áu ō
*ei
ei ī ei /ī/ ē
*eu
eu ēo iu ō
*oi
oi ā ái ē
*ou
ou ēa áu ō
*p
p f f; b p b in Gothic by Verner's law
*t
t þ/ð; d þ; d t þ and ð are different graphs for the same sound; d in the Germanic languages by Verner's law
*ḱ
k h; g h; g ś g in the Germanic languages by Verner's law
*k
k; c (+ PIE e/i)
*kʷ
p; t (+ e/i) hw, h; g, w ƕ /hʷ/; g, w, gw g, w, gw in the Germanic languages by Verner's law
*b
b p p b
*d
d t t d
*ǵ
g k k j
*g
g; j (+ PIE e/i)
*gʷ
b; d (+ i) q, c q
*bʰ
ph; p b b bh; b Greek p, Sanskrit b before any aspirated consonant (Grassmann's law)
*dʰ
th; t d d dh; d Greek t, Sanskrit d before any aspirated consonant
*ǵʰ
kh; k g g h; j Greek k, Sanskrit j before any aspirated consonant
*gʰ
gh; g
h; j (+ PIE e/i) Greek k, Sanskrit g, j before any aspirated consonant
*gʷʰ
ph; p
th; t (+ e/i) b (word-initially);
g, w b (word-initially);
g, w, gw Greek p, t, Sanskrit g, j before any aspirated consonant
*s
h (word-initially); s, - s; r s; z s, ṣ r, z in Germanic by Verner's law
*y
h, z (word-initially); - g(e) /j/ j /j/ y
*w
- w w v
Pure vowels[edit]
Initial syllables[edit]
In initial syllables, Latin generally preserves all of the simple vowels of Proto-Italic (see above):
PIE *h₂eǵros "field" > *agro- > ager, gen. agrī (cf. Greek agrós, English acre, Sanskrit ájra-)
PIE *kápr̥- "penis" > *kapros > caper "he-goat", gen. caprī (cf. Greek kápros "boar", Old English hæfer "he-goat", Sanskrit kápr̥th- "penis")
PIE *seḱs "six", septḿ̥ "seven" > sex, septem (cf. Greek heks, heptá, Lithuanian šešì, septynì, Sanskrit ṣáṣ, saptá)
PIE *kʷis > quis "who?" (Greek tís,[6] Avestan čiš, Sanskrit kíḥ)
PIE *kʷod > quod "what, that" (relative) (Old English hwæt "what", Sanskrit kád)
PIE *oḱtōu "eight" > octō (Greek oktṓ, Irish ocht, Sanskrit aṣṭā́(u))
PIE *nokʷts "night" > nox, gen. noctis (Greek nuks < *nokʷs, Sanskrit nák < *nakts, Lithuanian naktìs)
PIE *(H)i̯ug-óm "yoke" > iugum (Greek zugón, Gothic juk, Sanskrit yugá-)
PIE *méh₂tēr "mother" > *mā́tēr > māter (Doric Greek mā́tēr, Old Irish máthir, Sanskrit mātár-)
PIE *sweh₂dús "pleasing, tasty" > *swādu- > *swādwi- (remade into i-stem) > suāvis (Doric Greek hādús, English sweet, Sanskrit svādú-)
PIE *sēmi- (or *seh₁mi-) "half" > sēmi- (Greek hēmi-, Old English sām-)
PIE *gneh₃-tó- "known" > *gnō-tó- > nōtus (cf. i-gnōtus "unknown"; Welsh gnawd "customary", Sanskrit jñātá-; Greek gnōtós[7])
PIE *muHs "mouse" > mūs (cf. Old English mūs, Greek mûs, Sanskrit mū́ṣ-)
Short vowel changes in initial syllables:
*swe- > so-:
*swesōr > soror, gen. sorōris "sister"
*swépnos (or *swópnos) > somnus "sleep"
*we- > wo- before labial consonants or velarized l [ɫ] (l pinguis; i.e. an l not followed by i, ī or l):
*welō "I want" > volō (vs. velle "to want" before l exīlis)
*wemō "I vomit" > vomō (cf. Greek eméō, Sanskrit vámiti)
e > i before [ŋ] (spelled n before a velar, or g before n):
PIE *dn̥ǵʰwéh₂s > *dn̥ɣwā > *denɣʷā > Old Latin dingua > lingua "tongue" (l- from lingō "to lick")
PIE *pénkwe "five" > *kʷenkʷe > quīnque (long ī from quīntus "fifth" < *quīnctos, where lengthening is normal before nct)
PIE *deḱ-no- > *deŋno- > dignus "worthy"
There are numerous examples where PIE *o appears to result in Latin a instead of expected o, mostly next to labial or labializing consonants. A group of cases showing *-ow- > -av- (before stress), *-ōw- > -āv- is known as Thurneysen-Havet's law[8]: examples include PIE *lowh₃ṓ > *lowṓ > lavō 'I wash'; PIE *oḱtōwos > octāvus 'eighth' (but octō 'eight'). Other cases remain more disputed, such as mare 'sea', in contrast to Irish muir, Welsh môr (Proto-Celtic *mori) < PIE *móri; lacus 'lake', in contrast to Irish loch < PIE *lok-u-. De Vaan (2008: 8) suggests a general shift *o > a in open syllables when preceded by any of *b, *m; *kʷ, *w; *l. Vine (2011)[9] disputes the cases with *moCV, but proposes inversely that *mo- > ma- when followed by r plus a velar (k or g).
Medial syllables[edit]
In non-initial syllables, there was more vowel reduction of short vowels. The most extreme case occurs with short vowels in medial syllables (i.e. short vowels in a syllable that is neither the first nor the last), where all five vowels usually merge into a single vowel:
1. They merge into e before r (sometimes original o is unaffected)
*in-armis > inermis "unarmed" (vs. arma "arms")
Latin-Faliscan *pe-par-ai "I gave birth" > peperī (vs. pariō "I give birth")
*kom-gesō > congerō "to collect" (vs. gerō "to do, carry out")
*kinis-es "ash" (gen.sg.) > cineris (vs. nom.sg. cinis)
*Falisioi > Faleriī "Falerii (major town of the Faliscans)" (vs. Faliscus "Faliscan")
*-foro- "carrying" (cf. Greek -phóros) > -fero-, e.g. furcifer "gallows bird"
PIE *swéḱuros "father-in-law" > *swekuros > Old Latin *soceros > socer, gen. socerī
2. They become Old Latin o > u before l pinguis, i.e., an l not followed by i, ī, or l:
*en-saltō "to leap upon" > īnsoltō (with lengthening before ns) > īnsultō (vs. saltō "I leap")
*ad-alēskō "to grow up" > adolēscō > adulēscō (vs. alō "I nourish")
*ob-kelō "to conceal" > occulō (vs. celō "I hide")
Greek Sikelós "a Sicilian" > *Sikolos > Siculus (vs. Sicilia "Sicily")
te-tol-ai > tetulī "I carried" (formerly l pinguis here because of the original final -ai)
kom-solō "deliberate" > cōnsulō
PIE *-kl̥d-to- "beaten" > *-kolsso-[10] > perculsus "beaten down"
3. But they remain o before l pinguis when immediately following a vowel:
Latin-Faliscan *fili-olos > filiolus "little son"
Similarly, alveolus "trough"
4. Before /w/ the result is always u, in which case the /w/ is not written:
*eks-lawō "I wash away" > ēluō
*mon-i-wai "I warned" > monuī
*tris-diw-om "period of three days" > trīduom > trīduum
*dē nowōd "anew" > dēnuō
5. They become i before one consonant other than r or l pinguis:
*re-fakjō "to remake" > *refakiō > reficiō (vs. faciō "I do, make")
Latin-Faliscan *ke-kad-ai "I fell" > cecidī (vs. cadō "I fall")
*ad-tenējō > attineō "to concern" (vs. teneō "I hold")
*kom-regō > corrigō "to set right, correct" (vs. regō "I rule; straighten")
Greek Sikelía "Sicily" > Sicilia (vs. Siculus "a Sicilian")
PIE *me-món-h₂e (perfect) "thought, pondered" > Latin-Faliscan *me-mon-ai > meminī "I remember"
*kom-itājō "accompany" > comitō
*nowotāts "newness" > novitās
*kornu-kan- "trumpeter" > cornicen
*kaput-es "head" (gen. sg.) > capitis (vs. nom.sg. caput)
6. But they sometimes become e before one consonant other than r or l pinguis, when immediately following a vowel:
*sokiotāts "fellowship" > societās
*wariogājesi "to make diverse" > variegāre
But: *tībia-kan- "flute-player" > *tībiikan- > tībīcen
But: *medio-diēs "midday" > *meriodiēs (dissimilative rhotacism) > *meriidiēs > merīdiēs "noon; south"
7. Variation between i and (often earlier) u is common before a single labial consonant (p, b, f, m), underlyingly the sonus medius vowel:
From the root *-kap- "grab, catch":
occupō "seize" vs. occipiō "begin"
From the related noun *-kaps "catcher": prīnceps "chief" (lit. "seizer of the first (position)"), gen. prīncipis, vs. auceps "bird catcher", gen. aucupis
*man-kapiom > mancupium "purchase", later mancipium
*sub-rapuit > surrupuit "filches", later surripuit
*mag-is-emos > maxumus "biggest", later maximus; similarly proxumus "nearest", optumus "best" vs. later proximus, optimus
*pot-s-omos > possumus "we can"; *vel-omos > volumus "we want"; but *leg-omos > legimus "we gather", and all other such verbs (-umus is isolated in sumus, possumus and volumus)
Medially before two consonants, when the first is not r or l pinguis, the vowels do not merge to the same degree:
1. Original a, e and u merge into e:
Greek tálanton > *talantom > talentum
*sub-raptos "filched" > surreptus (vs. raptus "seized")
*re-faktos "remade" > refectus (cf. factus "made")
*ad-tentos > attentus "concerned" (cf. tentus "held", attineō "to concern")
2. But original i is unaffected:
*re-likʷtos "left (behind)" > relictus
3. And original o raises to u:
*legontor "they gather" > leguntur
*ejontes "going" (gen. sg.) > euntis
rōbos-to- > rōbustus "oaken" (cf. rōbur "oak" < *rōbos)
Syncope[edit]
Exon's Law dictates that if there are two light medial syllables in a row (schematically, σσ̆σ̆σ, where σ = syllable and σ̆ = light syllable, where "light" means a short vowel followed by only a single consonant), the first syllable syncopates (i.e. the vowel is deleted):
*deksiteros "right (hand)" > dexterus (cf. Greek deksiterós)
*re-peparai > repperī "I found" (cf. peperī "I gave birth" < *peparai)
*prīsmo-kapes > prīncipis "prince" gen. sg. (nom. sg. prīnceps < *prīsmo-kaps by analogy)
*mag-is-emos > maximus "biggest" (cf. magis "more")
Syncopation tends to occur after r and l in all non-initial syllables, sometimes even in initial syllables.[11]
*feret "he carries" > fert
*agros "field" > *agr̩s > *agers > *agerr > ager
*imbris "rainstorm" > *imbers > imber
*tris "three times" > *tr̩s > *ters > Old Latin terr > ter
*faklitāts > facultās
Sometimes early syncope causes apparent violations of Exon's Law:
kosolinos "of hazel" > *kozolnos (not **koslinos) > *korolnos > *korulnos (o > u before l pinguis, see above) > colurnus (metathesis)
Syncope of -i- also occurred in -ndis, -ntis and -rtis.[11] -nts then became -ns with lengthening of the preceding vowel, while -rts was simplified to -rs without lengthening.
*montis "hill" > *monts > mōns
*gentis "tribe" > *gents > gēns
*frondis "leaf" > *fronts > frōns
*partis "part" > *parts > pars
Final syllables[edit]
In final syllables of polysyllabic words before a final consonant or cluster, short a, e, i merge into either e or i depending on the following consonant, and short o, u merge into u.
1. Short a, e, i merge into i before a single non-nasal consonant:
Proto-Italic *rededas, *rededat > reddis, reddit "you return, he returns"
PIE thematic 2nd/3rd sg. *-esi, *-eti > *-es, *-et > -is, -it (e.g. legis, legit "you gather, he gathers")
i-stem nom. sg. *-is > -is
2. Short a, e, i merge into e before a cluster or a single nasal consonant:
*prīsmo-kaps > prīnceps "first, chief" (cf. capiō "to take")
*kornu-kan-(?s) > cornicen "trumpeter" (cf. canō "to sing")
*mīlets > mīles "soldier"
*septḿ̥ > septem "seven"
i-stem acc. sg. *-im > em
3. Short o, u merge into u:
o-stem nominative *-os > Old Latin -os > -us
o-stem accusative *-om > Old Latin -om > -um
PIE thematic 3rd pl. *-onti > *-ont > -unt
PIE *i̯ekʷr̥ > *jekʷor > iecur "liver"
PIE thematic 3rd sg. mediopassive *-etor > -itur
*kaput > caput "head"
4. All short vowels apparently merge into -e in absolute final position.[dubious – discuss]
Long vowels in final syllables shorten before most consonants (but not final s), yielding apparent exceptions to the above rules:
Proto-Italic *amāt > amat "he/she loves" (cf. passive amātur)
Proto-Italic *amānt > amant "they love"
a-stem acc. sg. *-ām > -am
PIE thematic 1st sg. mediopassive *-ōr > -or
*swesōr > soror "sister" (cf. gen. sorōris)
Diphthongs[edit]
Initial syllables[edit]
Proto-Italic maintained all PIE diphthongs except for the change *eu > *ou. The Proto-Italic diphthongs tend to remain into Old Latin but generally reduce to pure long vowels by Classical Latin.
1. PIE *ei > Old Latin ei > ẹ̄, a vowel higher than ē < PIE *ē. This then developed to ī normally, but to ē before v:
PIE *deiḱ- "point (out)" > Old Latin deicō > dīcō "to say"
PIE *bʰeydʰ- "be persuaded, be confident" > *feiðe- > fīdō "to trust"
PIE *deywós "god, deity" > Very Old Latin deiuos (Duenos inscription) > dẹ̄vos > deus
But nominative plural *deivoi > *deivei > *dẹ̄vẹ̄ > dīvī > diī; vocative singular *deive > *dẹ̄ve > dīve
2. PIE *eu, *ou > Proto-Italic *ou > Old Latin ou > ọ̄ (higher than ō < PIE *ō) > ū:
PIE *(H)jeug- "join" > *joug-s-mn-to- > Old Latin iouxmentom "pack horse" > iūmentum
PIE *louk-s-neh₂ > *louksnā > Old Latin losna (i.e. lọ̄sna) > lūna "moon" (cf. Old Prussian lauxnos "stars", Avestan raoχšnā "lantern")
PIE *deuk- > *douk-e- > Old Latin doucō > dūcō "lead"
3. PIE *ai, *Hei > ae:
PIE *kh₂ei-ko- > *kaiko- > caecus "blind" (cf. Old Irish cáech /kaiχ/ "blind", Gothic háihs "one-eyed", Sanskrit kekara- "squinting")
4. PIE *au > au:
PIE *h₂eug- > *augeje/o > augeō "to increase" (cf. Greek aúksō, Gothic áukan, Lithuanian áugti)
5. PIE *oi > Old Latin oi, oe > ū (occasionally preserved as oe):
PIE *h₁oi-nos > Old Latin oinos > oenus > ūnus "one"
Greek Phoiniks > Pūnicus "Phoenician"
But: PIE *bʰoidʰ- > *foiðo- > foedus "treaty" (cf. fīdō above)
Medial syllables[edit]
All diphthongs in medial syllables become ī or ū.
1. (Post-)PIE *ei > ī, just as in initial syllables:
*en-deik-ō > indīcō "to point out" (cf. dīcō "to say")
2. (Post-)PIE *oi > ū, just as in initial syllables:
PIE *n̥-poini "with impunity" > impūne (cf. poena "punishment").
3. (Post-)PIE *eu, *ou > Proto-Italic *ou > ū, just as in initial syllables:
*en-deuk-ō > *indoucō > indūcō "to draw over, cover" (cf. dūcō "to lead")
4. Post-PIE *ai > Old Latin ei > ī:
*ke-kaid-ai "I cut", perf. > cecīdī (cf. caedō "I cut", pres.)
*en-kaid-ō "cut into" > incīdō (cf. caedō "cut")
Early Greek (or from an earlier source) *elaíwā "olive" > olīva
5. Post-PIE *au > ū (rarely oe):
*en-klaud-ō "enclose" > inclūdō (cf. claudō "close")
*ad-kauss-ō "accuse" > accūsō (cf. causa "cause")
*ob-aud-iō "obey" > oboediō (cf. audiō "hear")
Final syllables[edit]
Mostly like medial syllables:
*-ei > ī: PIE *meh₂tr-ei "to mother" > mātrī
*-ai > ī in multisyllabic words: Latin-Faliscan peparai "I brought forth" > peperī
*-eu/ou- > ū: post-PIE manous "hand", gen. sg. > manūs
Different from medial syllables:
-oi > Old Latin -ei > ī (not ū): PIE o-stem plural *-oi > -ī (cf. Greek -oi);
-oi > ī also in monosyllables: PIE kʷoi "who" > quī
-ai > ae in monosyllables: PIE *prh₂ei "before" > prae (cf. Greek paraí)
Syllabic resonants and laryngeals[edit]
The PIE syllabic resonants *m̥, *n̥, *r̥, *l̥ generally become em, en, or, ol (cf. Greek am/a, an/a, ar/ra, al/la; Germanic um, un, ur, ul; Sanskrit am/a, an/a, r̥, r̥; Lithuanian im̃, iñ, ir̃, il̃):
PIE *déḱm̥(t) "ten" > decem (cf. Irish deich, Greek deka, Gothic taíhun /tɛhun/)
PIE *(d)ḱm̥tóm "hundred" > centum (cf. Welsh cant, Gothic hund, Lithuanian šim̃tas)
PIE *n̥- "not" > OL en- > in- (cf. Greek a-/an-, English un-)
PIE *tn̥tós "stretched" > tentus (cf. Greek tatós, Sanskrit tatá-)
PIE *ḱr̥d- "heart" > *cord > cor (cf. Greek kēr, English heart, Lithuanian širdìs)
PIE *ml̥dú- "soft" > *moldu- > *mollwi- (remade as i-stem) > *molwi- > mollis (cf. Irish meldach "pleasing", English mild, Czech mladý)
The laryngeals *h₁, *h₂, *h₃ appear in Latin as a when between consonants, as in most languages (but Greek e/a/o respectively, Sanskrit i):
PIE *dʰh₁-tós "put" > L factus, with /k/ of disputed etymology (cf. Greek thetós, Sanskrit hitá- < *dhitá-)
PIE *ph₂tḗr "father" > L pater (cf. Greek patḗr, Sanskrit pitā́, English father)
PIE *dh₃-tós "given" > L datus (cf. Greek dotós, Sanskrit ditá-)
A sequence of syllabic resonant + laryngeal, when before a consonant, produced mā, nā, rā, lā (cf. Greek nē/nā/nō, rē/rā/rō, etc. depending on the laryngeal; Germanic um, un, ur, ul; Sanskrit ā, ā, īr/ūr, īr/ūr; Lithuanian ím, ín, ír, íl):
PIE *ǵr̥h₂-no- "grain" > grānum (cf. Old Irish grán, English corn, Lithuanian žìrnis "pea")
PIE *h₂wl̥h₁-neh₂ "wool" > *wlānā > lāna (cf. Welsh gwlân, Greek lēnos, Lithuanian vìlna)
PIE *ǵn̥h₁-to- "born" > gnātus "son" (cf. Middle Welsh gnawt "relative", Greek dió-gnētos "Zeus' offspring", English kind)
Consonants[edit]
Aspirates[edit]
The Indo-European voiced aspirates bʰ, dʰ, gʰ, gʷʰ, which were probably breathy voiced stops, first devoiced in initial position (fortition), then fricatized in all positions, producing pairs of voiceless/voiced fricatives in Proto-Italic: f ~ β, θ ~ ð, χ ~ ɣ, χʷ ~ ɣʷ respectively.[12] The fricatives were voiceless in initial position. However, between vowels and other voiced sounds, there are indications — in particular, their evolution in Latin — that the sounds were actually voiced. Likewise, Proto-Italic /s/ apparently had a voiced allophone [z] in the same position.
In all Italic languages, the word-initial voiceless fricatives f, θ, and χʷ all merged to f; thus, in Latin, the normal outcome of initial PIE bʰ, dʰ, gʰ, gʷʰ is f, f, h, f, respectively. Examples:
PIE *bʰréh₂tēr "brother" > *bʰrā́tēr > frāter (cf. Old Irish bráthair, Sanskrit bhrā́tar-, Greek phrā́tēr "member of a phratry")
PIE *bʰére "carry" > ferō (cf. Old Irish beirim "I bear", English bear, Sanskrit bhárati)
PIE *dʰwṓr "door" > θwor- > *forā > forēs (pl.) "door(s)" (cf. Welsh dôr, Greek thurā, Sanskrit dhvā́raḥ (pl.))
PIE *dʰeh₁- "put, place" > *dʰh₁-k- > *θaki- > faciō "do, make" (cf. Welsh dodi, English do, Greek títhēmi "I put", Sanskrit dádhāti he puts")
PIE *gʰabʰ- "seize, take" > *χaβ-ē- > habeō "have" (cf. Old Irish gaibid "takes", Old English gifan "to give", Polish gabać "to seize")
PIE *ǵʰh₂ens "goose" > *χans- > (h)ānser (cf. Old Irish géiss "swan", German Gans, Greek khḗn)
PIE *ǵʰaidos "goat" > *χaidos > hædus "kid" (cf. Old English gāt "goat", Polish zając "hare", Sanskrit háyas "horse")
PIE *gʷʰerm- "warm" > *χʷormo- > formus (cf. Old Prussian gorme "heat", Greek thermós, Sanskrit gharmáḥ "heat")
PIE *gʷʰen-dʰ- "to strike, kill" > *χʷ(e)nð- > fendō (cf. Welsh gwanu "to stab", Old High German gundo "battle", Sanskrit hánti "(he) strikes, kills")
Word-internal *-bʰ-, *-dʰ-, *-gʰ-, *-gʷʰ- evolved into Proto-Italic β, ð, ɣ, ɣʷ. In Osco-Umbrian, the same type of merger occurred as that affecting voiceless fricatives, with β, ð, and ɣʷ merging to β. In Latin, this did not happen, and instead the fricatives defricatized, giving b, d ~ b, g ~ h, g ~ v ~ gu.
*-bʰ- is the simplest case, consistently becoming b.
PIE *bʰébʰrus "beaver" > *feβro > Old Latin feber > fiber
*-dʰ- usually becomes d, but becomes b next to r or u, or before l.
PIE *bʰeidʰ- "be persuaded" > *feiðe > fīdō "I trust" (cf. Old English bīdan "to wait", Greek peíthō "I trust")
PIE *medʰi-o- "middle" > *meðio- > medius (cf. Old Irish mide, Gothic midjis, Sanskrit mádhya-)
PIE *h₁rudʰ-ró- "red" > *ruðro- > ruber (cf. Old Russian rodrŭ, Greek eruthrós, Sanskrit rudhirá-)
PIE *werh₁-dʰh₁-o- "word" > *werðo- > verbum (cf. English word, Lithuanian var̃das)
PIE *sth̥₂-dʰlom > *staðlom > stabulum "abode" (cf. German Stadel)
PIE *krei(H)-dʰrom "sieve, sifter" > *kreiðrom > crībrum "sieve" (cf. Old Irish críathar /krīǝ̯θǝρ/ "sieve", Old English hrīder "sieve")
The development of *-gʰ- is twofold: *-gʰ- becomes h [ɦ] between vowels but g elsewhere:
PIE *weǵʰ- "carry" > *weɣ-e/o > vehō (cf. Greek okhéomai "I ride", Old English wegan "to carry", Sanskrit váhati "(he) drives")
PIE *dʰi-n-ǵʰ- "shapes, forms" > *θinɣ-e/o > fingō (cf. Old Irish -ding "erects, builds", Gothic digan "to mold, shape")
*-gʷʰ- has three outcomes, becoming gu after n, v between vowels, and g next to other consonants. All three variants are visible in the same root *snigʷʰ- "snow" (cf. Irish snigid "snows", Greek nípha):
PIE *snigʷʰ-s > *sniɣʷs > *nigs > nom. sg. nix "snow"
PIE *snigʷʰ-ós > *sniɣʷos > *niβis > gen. sg. nivis "of snow"
PIE *snei-gʷʰ-e/o > *sninɣʷ-e/o (with n-infix) > ninguit "it snows"
Other examples:
PIE *h₁le(n)gʷʰu- > *h₁legʷʰu- > *leɣʷus > *leβwi- (remade as i-stem) > levis "lightweight" (cf. Welsh llaw "small, low", Greek elakhús "small", Sanskrit laghú-, raghú- "quick, light, small")
Labiovelars[edit]
*gʷ has results much like non-initial *-gʷʰ, becoming v /w/ in most circumstances, but gu after a nasal and g next to other consonants:
PIE *gʷih₃wos > *ɣʷīwos > vīvus "alive" (cf. Old Irish biu, beo, Lithuanian gývas, Sanskrit jīvá- "alive")
PIE *gʷm̥i̯e/o- "come" > *ɣʷen-je/o > veniō (cf. English come, Greek baínō "I go", Avestan ǰamaiti "he goes")
PIE *gʷr̥h₂us "heavy" > *ɣʷraus > grāvis (cf. Greek barús, Gothic kaúrus, Sanskrit gurú-)
PIE *h₃engʷ- > *onɣʷ-en > unguen "salve" (cf. Old Irish imb "butter", Old High German ancho "butter", Sanskrit añjanam "anointing, ointment")
PIE *n̥gʷén- "(swollen) gland" > *enɣʷen > inguen "bubo; groin" (cf. Greek adḗn gen. adénos "gland", Old High German ankweiz "pustules")
*kʷ remains as qu before a vowel, but reduces to c /k/ before a consonant or next to a u:
PIE *kʷetwóres, neut. *kʷetwṓr "four" > quattuor (cf. Old Irish cethair, Lithuanian keturì, Sanskrit catvā́raḥ)
PIE *sekʷ- "to follow" > sequor (cf. Old Irish sechem, Greek hépomai, Sanskrit sácate)
PIE *leikʷ- (pres. *li-né-kʷ-) "leave behind" > *linkʷ-e/o- : *likʷ-ē- > linquō "leaves" : liceō "is allowed; is for sale" (cf. Greek leípō, limpánō, Sanskrit riṇákti, Gothic leiƕan "to lend")
PIE *nokʷts "night" > nox, gen. sg. noctis
The sequence *p ... *kʷ assimilates to *kʷ ... *kʷ:
PIE *pénkʷe "five" > quīnque (cf. Old Irish cóic, Greek pénte, Sanskrit páñca)
PIE *pérkʷus "oak" > quercus (cf. Trentino porca "fir", Punjabi pargāī "holm oak", Gothic faírƕus "world", faírgun- "mountain"[13])
PIE *pekʷō "I cook" > *kʷekʷō > coquō (cf. coquīna, cocīnā "kitchen" vs. popīna "tavern" < Oscan, where *kʷ > p)
The sequences *ḱw, *ǵw, *ǵʰw develop identically to *kʷ, *gʷ, *gʷʰ:
PIE *éḱwos "horse" > *ekʷos > Old Latin equos > ecus > equus (assimilated from other forms, e.g. gen. sg. equī; cf. Sanskrit aśva-, which indicates -ḱw- not -kʷ-)
PIE *ǵʰweh₁ro- "wild animal" > *χʷero- > ferus (cf. Greek thḗr, Lesbian phḗr, Lithuanian žvėrìs)
PIE *mreǵʰus "short" > *mreɣu- > *mreɣwi- (remade as i-stem) > brevis (cf. Old English myrge "briefly", Greek brakhús, Avestan mǝrǝzu-)
PIE *dn̥ǵʰwéh₂[14] "tongue" > *dn̥ɣwā > *denɣʷā > Old Latin dingua > lingua
Other sequences[edit]
Initial *dw- (attested in Old Latin as du-) becomes b-, thus compensating for the dearth of words beginning with *b in PIE:
PIE *dwis "twice" > duis > bis (cf. Greek dís, Sanskrit dvis)
PIE *deu-l̥- "injure" > duellom "war" > bellum (a variant duellum survived in poetry as a trisyllabic word, whence English "duel")
S-rhotacism[edit]
Indo-European s between vowels was first voiced to [z] in late Proto-Italic and became r in Latin and Umbrian, a change known as rhotacism. Early Old Latin documents still have s [z], and Cicero once remarked that a certain Papirius Crassus officially changed his name from Papisius in 339 b.c.,[15] indicating the approximate time of this change. This produces many alternations in Latin declension:
flōs "flower", gen. flōris
mūs "mouse", pl. mūrēs
est "he is", fut. erit "he will be"
Other examples:
Proto-Italic *ausōs, ausōsem > *auzōs, auzōzem > aurōra "dawn" (change of suffix; cf. English east, Aeolic Greek aúōs, Sanskrit uṣā́s)
Proto-Italic *swesōr > *swozōr > soror "sister" (cf. Old English sweostor, Sanskrit svásar)
However, before another r, dissimilation occurred with sr [zr] becoming br (likely via an intermediate *ðr):
Proto-Italic *swesr-īnos > *swezrīnos ~ *sweðrīnos > sobrīnus "maternal cousin" (with so- by analogy to soror)
Proto-Italic *keras-rom > *kerazrom ~ *keraðrom > cerebrum "skull, brain" (cf. Greek kéras "horn")
See also[edit]
De vulgari eloquentia
Legacy of the Roman Empire
Notes[edit]
Jump up ^ Leonard Robert Palmer - The Latin language - 372 pages University of Oklahoma Press, 1987 Retrieved 2012-02-01 ISBN 0-8061-2136-X
Jump up ^ Ramat, Anna G.; Paolo Ramat (1998). The Indo-European Languages. Routledge. pp. 272–75. ISBN 0-415-06449-X.
Jump up ^ Ramat, Anna G.; Paolo Ramat (1998). The Indo-European Languages. Routledge. p. 313. ISBN 0-415-06449-X.
Jump up ^ Timothy J. Pulju Rice University .edu/~ Retrieved 2012-02-01
Jump up ^ Allen, W. Sidney (1989). Vox Latina. Cambridge University Press. pp. 83–84. ISBN 0-521-22049-1.
Jump up ^ kʷi- > ti- is normal in Attic Greek; Thessalian Greek had kís while Cypro-Arcadian had sís.
Jump up ^ Greek is ambiguously either < *gneh₃-tó- or *gn̥h₃-tó-
Jump up ^ Collinge, N. E. (1985). The Laws of Indo-European. John Benjamins. pp. 193–195. ISBN 90-272-3530-9.
Jump up ^ Vine, Brent (2011). "Initial *mo- in Latin and Italic". Münchener Studien zur Sprachwissenschaft (65): 261–286.
Jump up ^ l̥ > ol is normal in Proto-Italic.
^ Jump up to: a b Sihler, New Comparative Grammar of Greek and Latin, 1995
Jump up ^ James Clackson & Geoffrey Horrocks, The Blackwell History of the Latin Language (Oxford: Blackwell, 2007), 51-2.
Jump up ^ Both "world" and "mountain" evolve out of the early association of oak trees with strength, cf. Latin robur = "oak" but also "strength"
Jump up ^ PIE *dn̥ǵhwéh₂; -ǵʰw- not -gʷʰ- indicated by Old Church Slavonic języ-kŭ "tongue" < *n̥ǵhu-H-k- with loss of initial *d-; -gʷh- would yield /g/, not /z/.
Jump up ^ Fortson, Benjamin W., Indo-European Language and Culture: An Introduction, p. 283
Sources[edit]
Allen, J. H.; James B. Greenough (1931). New Latin Grammar. Boston: Ginn and Company. ISBN 1-58510-027-7.
External links[edit]
Latin Etymology, An Etymological Dictionary of the Latin Language
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Cortical homunculus
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A 2-D cortical sensory homunculus
A cortical homunculus is a distorted representation of the human body, based on a neurological "map" of the areas and proportions of the brain dedicated to processing motor functions, or sensory functions, for different parts of the body. Homunculus is Latin for "little man", and was a term used in alchemy and folklore prior to the concept being utilized in scientific literature. A cortical homunculus, or "cortex man", illustrates the concept of a representation of the body lying within the brain.
Contents
[hide]
1 Types
2 Arrangement
3 Discovery
4 Representation
5 See also
6 References
7 External links
Types[edit]
A 2-D cortical motor homunculus
A motor homunculus represents a map of brain areas dedicated to motor processing for different anatomical divisions of the body. The primary motor cortex is located in the precentral gyrus, and handles signals coming from the premotor area of the frontal lobes.[1]
A sensory homunculus represents a map of brain areas dedicated to sensory processing for different anatomical divisions of the body. The primary sensory cortex is located in the postcentral gyrus, and handles signals coming from the thalamus.[1]
These signals are transmitted on from the gyri to the brain stem and spinal cord via corresponding nerves.
Arrangement[edit]
Along the length of the primary motor and sensory cortices, the areas specializing in different parts of the body are arranged in an orderly manner, although ordered differently than one might expect. The toes are represented at the top of the cerebral hemisphere (or more accurately, "the upper end", since the cortex curls inwards and down at the top), and then as one moves down the hemisphere, progressively higher parts of the body are represented, assuming a body that's faceless and has arms raised. Going further down the cortex, the different areas of the face are represented, in approximately top-to-bottom order, rather than bottom-to-top as before. The homunculus is split in half, with motor and sensory representations for the left side of the body on the right side of the brain, and vice-versa.[2]
The amount of cortex devoted to any given body region is not proportional to that body region's surface area or volume, but rather to how richly innervated that region is. Areas of the body with more complex and/or more numerous sensory or motor connections are represented as larger in the homunculus, while those with less complex and/or less numerous connections are represented as smaller. The resulting image is that of a distorted human body, with disproportionately huge hands, lips, and face.
In the sensory homunculus, below the areas handling sensation for the teeth, gums, jaw, tongue, and pharynx lies an area for intra-abdominal sensation. At the very top end of the primary sensory cortex, beyond the area for the toes, it has traditionally been believed that the sensory neural networks for the genitals occur. However, more recent research has suggested that there may be two different cortical areas for the genitals, possibly differentiated by one dealing with erogenous stimulation and the other dealing with non-erogenous stimulation.[3][4][5]
Discovery[edit]
3-D Sensory and Motor homunculus models at the Natural History Museum, London
Dr. Wilder Penfield and his co-investigators Edwin Boldrey and Theodore Rasmussen are considered to be the originators of the sensory and motor homunculi. They were not the first scientists to attempt to objectify human brain function by means of a homunculus.[5] However, they were the first to differentiate between sensory and motor function and to map the two across the brain separately, resulting in two different homunculi. In addition, their drawings and later drawings derived from theirs became perhaps the most famous conceptual maps in modern neuroscience because they compellingly illustrated the data at a single glance.[5]
Penfield first conceived of his homunculi as a thought experiment, and went so far as to envision an imaginary world in which the homunculi lived, which he referred to as "if". He and his colleagues went on to experiment with electrical stimulation of different brain areas of patients undergoing open brain surgery to control epilepsy, and were thus able to produce the topographical brain maps and their corresponding homunculi.[5][6]
More recent studies have improved this understanding of somatotopic arrangement using techniques such as functional magnetic resonance imaging (fMRI).[7]
Representation[edit]
Penfield referred to his creations as "grotesque creatures" due to their strange-looking proportions. For example, the sensory nerves arriving from the hands terminate over large areas of the brain, resulting in the hands of the homunculus being correspondingly large. In contrast, the nerves emanating from the torso or arms cover a much smaller area, thus the torso and arms of the homunculus look comparatively small and weak.
Penfield's homunculi are usually shown as 2-D diagrams. This is an oversimplification, as it cannot fully show the data set Penfield collected from his brain surgery patients. Rather than the sharp delineation between different body areas shown in the drawings, there is actually significant overlap between neighboring regions. The simplification suggests that lesions of the motor cortex will give rise to specific deficits in specific muscles. However, this is a misconception, as lesions produce deficits in groups of synergistic muscles. This finding suggests that the motor cortex functions in terms of overall movements as coordinated groups of individual motions.
The sensorimotor homunculi can also be represented as 3-D figures (such as the sensory homunculus sculpted by Sharon Price-James shown from different angles below), which can make it easier for laymen to understand the ratios between the different body regions' levels of motor or sensory innervation. However, these 3-D models do not illustrate which areas of the brain are associated with which parts of the body.
See also[edit]
Homunculus
Somatotopic arrangement
Alice in Wonderland syndrome
References[edit]
^ Jump up to: a b Marieb, E., Hoehn, K. Human Anatomy and Physiology. 7th Ed. 2007. Pearson Benjamin Cummings: San Francisco.
Jump up ^ Saladin, Kenneth (2007). Anatomy and Physiology: The Unity of Form and Function. McGraw Hill. p. 544-546
Jump up ^ Covington, Jr., William Oates (2015-05-27). "Homunculus (Topographic) Diagram". willcov.com. Retrieved 2017-07-07.
Jump up ^ "The Neurocritic: A New Clitoral Homunculus?". 2009-08-10. Retrieved 2017-07-07.
^ Jump up to: a b c d Cazala, Fadwa; Vienney, Nicolas; Stoléru, Serge (2015-03-10). "The cortical sensory representation of genitalia in women and men: a systematic review". Socioaffective Neuroscience & Psychology. 5. doi:10.3402/snp.v5.26428. ISSN 2000-9011. PMC 4357265  . PMID 25766001. Retrieved 2017-07-07.
Jump up ^ Penfield, Wilder; Boldrey, Edwin (1937). "Somatic Motor And Sensory Representation In The Cerebral Cortex Of Man As Studied By Electrical Stimulation". Brain. 60 (4): 389-443. doi:10.1093/brain/60.4.389. Retrieved 26 March 2016.
Jump up ^ Grodd W, Hülsmann E, Lotze M, Wildgruber D, Erb M. Sensorimotor mapping of the human cerebellum: fMRI evidence of somatotopic organization. Hum Brain Mapp. 2001 Jun;13(2):55-73.
External links[edit]
Wikimedia Commons has media related to Cortical homunculus.
Mole-ratunculus – an analog of a sensory homunculus, except for a mole-rat, from the paper Somatosensory cortex dominated by the representation of teeth in the naked mole-rat brain
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