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Venus flytrap
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This article is about the plant. For other uses, see Venus Flytrap (disambiguation).
Venus flytrap
Leaf
Conservation status

Vulnerable (IUCN 2.3)[1]
Scientific classification
Kingdom: Plantae
Clade: Angiosperms
Clade: Eudicots
Order: Caryophyllales
Family: Droseraceae
Genus: Dionaea
Sol. ex J.Ellis 1768
Species: D. muscipula
Binomial name
Dionaea muscipula
J.Ellis
Distribution
Synonyms[2]
Dionea Raf., spelling variant
Dionaea corymbosa
(Raf.) Steud. (1840)
Dionaea crinita
Sol. (1990) as synonym
Dionaea dentata
D'Amato (1998) name published without description
Dionaea heterodoxa
D'Amato (1998) nom.nud.
Dionaea muscicapa
St.Hil. (1824) sphalm.typogr.
Dionaea sensitiva
Salisb. (1796)
Dionaea sessiliflora
(Raf.) Steud. (1840)
Dionaea uniflora
(Raf.) Steud. (1840)
Drosera corymbosa
Raf. (1833)
Drosera sessiliflora
Raf. (1833)
Drosera uniflora
Raf. (1833)
The Venus flytrap (also referred to as Venus's flytrap or Venus' flytrap), Dionaea muscipula, is a carnivorous plant native to subtropical wetlands on the East Coast of the United States in North Carolina and South Carolina.[3] It catches its prey—chiefly insects and arachnids—with a trapping structure formed by the terminal portion of each of the plant's leaves, which is triggered by tiny hairs on their inner surfaces. When an insect or spider crawling along the leaves contacts a hair, the trap prepares to close, snapping shut only if another contact occurs within approximately twenty seconds of the first strike. The requirement of redundant triggering in this mechanism serves as a safeguard against wasting energy by trapping objects with no nutritional value, and the plant will only begin digestion after five more stimuli to ensure it has caught a live bug worthy of consumption.
Dionaea is a monotypic genus closely related to the waterwheel plant (Aldrovanda vesiculosa) and sundews (Drosera), all of which belong to the family Droseraceae.
Contents
[hide]
1 Description
2 Etymology
3 Carnivory
3.1 Prey selectivity
3.2 Mechanism of trapping
3.3 Digestion
4 Evolution
4.1 Proposed evolutionary history
5 Habitat
6 Cultivation
6.1 Cultivars
7 Conservation
8 In alternative medicine
9 See also
10 References
11 External links
Description
In 1760, the North Carolina colonial governor, Arthur Dobbs, penned the first written description of the plant in a letter to English botanist Peter Collinson It was dated Brunswick, Jan. 24, 1760.[4][5][6]
“The great wonder of the vegetable kingdom is a very curious unknown species of Sensitive. It is a dwarf plant. The leaves are like a narrow segment of a sphere, consisting of two parts, like the cap of a spring purse, the concave part outwards, each of which falls back with indented edges (like an iron spring fox-trap); upon anything touching the leaves, or falling between them, they instantly close like a spring trap, and confine any insect or anything that falls between them. It bears a white flower. To this surprising plant I have given the name of Fly trap Sensitive.”
—  Arthur Dobbs
This seems to be the earliest notice of the plant and is before the letters of John Ellis on the subject.[7]
The Venus flytrap is a small plant whose structure can be described as a rosette of four to seven leaves, which arise from a short subterranean stem that is actually a bulb-like object. Each stem reaches a maximum size of about three to ten centimeters, depending on the time of year;[8] longer leaves with robust traps are usually formed after flowering. Flytraps that have more than seven leaves are colonies formed by rosettes that have divided beneath the ground.
Illustration from Curtis's Botanical Magazine by William Curtis (1746–1799)
The leaf blade is divided into two regions: a flat, heart-shaped photosynthesis-capable petiole, and a pair of terminal lobes hinged at the midrib, forming the trap which is the true leaf. The upper surface of these lobes contains red anthocyanin pigments and its edges secrete mucilage. The lobes exhibit rapid plant movements, snapping shut when stimulated by prey. The trapping mechanism is tripped when prey contacts one of the three hair-like trichomes that are found on the upper surface of each of the lobes. The mechanism is so highly specialized that it can distinguish between living prey and non-prey stimuli, such as falling raindrops;[9] two trigger hairs must be touched in succession within 20 seconds of each other or one hair touched twice in rapid succession,[9] whereupon the lobes of the trap will snap shut, typically in about one-tenth of a second.[10] The edges of the lobes are fringed by stiff hair-like protrusions or cilia, which mesh together and prevent large prey from escaping. These protrusions, and the trigger hairs (also known as sensitive hairs) are likely homologous with the tentacles found in this plant’s close relatives, the sundews. Scientists have concluded that the snap trap evolved from a fly-paper trap similar to that of Drosera.[11]
The holes in the meshwork allow small prey to escape, presumably because the benefit that would be obtained from them would be less than the cost of digesting them. If the prey is too small and escapes, the trap will usually reopen within 12 hours. If the prey moves around in the trap, it tightens and digestion begins more quickly.
Speed of closing can vary depending on the amount of humidity, light, size of prey, and general growing conditions. The speed with which traps close can be used as an indicator of a plant's general health. Venus flytraps are not as humidity-dependent as are some other carnivorous plants, such as Nepenthes, Cephalotus, most Heliamphora, and some Drosera.
The Venus flytrap exhibits variations in petiole shape and length and whether the leaf lies flat on the ground or extends up at an angle of about 40–60 degrees. The four major forms are: 'typica', the most common, with broad decumbent petioles; 'erecta', with leaves at a 45-degree angle; 'linearis', with narrow petioles and leaves at 45 degrees; and 'filiformis', with extremely narrow or linear petioles. Except for 'filiformis', all of these can be stages in leaf production of any plant depending on season (decumbent in summer versus short versus semi-erect in spring), length of photoperiod (long petioles in spring versus short in summer), and intensity of light (wide petioles in low light intensity versus narrow in brighter light).[citation needed]
Flowering venus flytrap showing its long flower stem
Closeup of flower (c. 20 mm in diameter)
The species produces small, shiny black seeds
Etymology
The plant's common name refers to Venus, the Roman goddess of love. The genus name, Dionaea ("daughter of Dione"), refers to the Greek goddess Aphrodite, while the species name, muscipula, is Latin for "mousetrap".[12]
Historically, the plant was also known by the slang term "tipitiwitchet" or "tippity twitchet", possibly an oblique reference to the plant's resemblance to human female genitalia.[12][13]
Carnivory
A closing trap
Prey selectivity
Play media
A time lapse showing Venus flytrap catching prey
Most carnivorous plants selectively feed on specific prey. This selection is due to the available prey and the type of trap used by the organism. With the Venus flytrap, prey is limited to beetles, spiders and other crawling arthropods. In fact, the Dionaea diet is 33% ants, 30% spiders, 10% beetles, and 10% grasshoppers, with fewer than 5% flying insects.[14] Given that Dionaea evolved from an ancestral form of Drosera (carnivorous plants that use a sticky trap instead of a snap trap) the reason for this evolutionary branching becomes clear. Whilst Drosera consume smaller, aerial insects, Dionaea consume larger terrestrial bugs. Dionaea are able to extract more nutrients from these larger bugs. This gives Dionaea an evolutionary advantage over their ancestral sticky trap form.[15]
Mechanism of trapping
Closeup of one of the hinged trigger hairs
The Venus flytrap is one of a very small group of plants capable of rapid movement, such as Mimosa pudica, the Telegraph plant, sundews and bladderworts.
The mechanism by which the trap snaps shut involves a complex interaction between elasticity, turgor and growth. The trap only shuts when there have been two stimulations of the trigger hairs; this is to avoid inadvertent triggering of the mechanism by dust and other wind-borne debris. In the open, untripped state, the lobes are convex (bent outwards), but in the closed state, the lobes are concave (forming a cavity). It is the rapid flipping of this bistable state that closes the trap,[10] but the mechanism by which this occurs is still poorly understood. When the trigger hairs are stimulated, an action potential (mostly involving calcium ions—see calcium in biology) is generated, which propagates across the lobes and stimulates cells in the lobes and in the midrib between them.[16] It is hypothesized that there is a threshold of ion buildup for the Venus flytrap to react to stimulation.[17] After closing, the flytrap counts additional stimulations of the trigger hairs, to five total, to start the production of digesting enzymes.[18] The acid growth theory states that individual cells in the outer layers of the lobes and midrib rapidly move 1H+ (hydrogen ions) into their cell walls, lowering the pH and loosening the extracellular components, which allows them to swell rapidly by osmosis, thus elongating and changing the shape of the trap lobe. Alternatively, cells in the inner layers of the lobes and midrib may rapidly secrete other ions, allowing water to follow by osmosis, and the cells to collapse. Both of these mechanisms may play a role and have some experimental evidence to support them.[19][20]
Digestion
If the prey is unable to escape, it will continue to stimulate the inner surface of the lobes, and this causes a further growth response that forces the edges of the lobes together, eventually sealing the trap hermetically and forming a "stomach" in which digestion occurs. Release of the digestive enzymes is controlled by the hormone jasmonic acid, the same hormone that triggers the release of toxins as an anti-herbivore defense mechanism in non-carnivorous plants. (See Evolution below)[18][21] Once the digestive glands in the leaf lobes have been activated, digestion is catalysed by hydrolase enzymes secreted by the glands.
Oxidative protein modification is likely to be a pre-digestive mechanism used by Dionaea muscipula. Aqueous leaf extracts have been found to contain quinones such as the naphthoquinone plumbagin that couples to different NADH-dependent diaphorases to produce superoxide and hydrogen peroxide upon autoxidation.[22] Such oxidative modification could rupture animal cell membranes. Plumbagin is known to induce apoptosis, associated with the regulation of the Bcl-2 family of proteins.[23] When the Dionaea extracts were pre-incubated with diaphorases and NADH in the presence of serum albumin (SA), subsequent tryptic digestion of SA was facilitated.[22] Since the secretory glands of Droseraceae contain proteases and possibly other degradative enzymes, it may be that the presence of oxygen-activating redox cofactors function as extracellular pre-digestive oxidants to render membrane-bound proteins of the prey (insects) more susceptible to proteolytic attacks.[22]
Digestion takes about ten days, after which the prey is reduced to a husk of chitin. The trap then reopens, and is ready for reuse.[24]
Evolution
Drosera falconeri, with short, wide, sticky leaf traps
Carnivory in plants is a very specialized form of foliar feeding, and is an adaptation found in several plants that grow in nutrient-poor soil. Carnivorous traps were naturally selected to allow these organisms to compensate for the nutrient deficiencies of their harsh environments by supplementing ordinary photosynthate with animal proteins.[25]
The "snap trap" mechanism characteristic of Dionaea is shared with only one other carnivorous plant genus, Aldrovanda. For most of the 20th century, this relationship was thought to be coincidental, more precisely an example of convergent evolution. Some phylogenetic studies even suggested that the closest living relatives of Aldrovanda were the sundews.[26] It was not until 2002 that a molecular evolutionary study, by analyzing combined nuclear and chloroplast DNA sequences, indicated that Dionaea and Aldrovanda were closely related and that the snap trap mechanism evolved only once in a common ancestor of the two genera.[27][28]
A 2009 study[26] presented evidence for the evolution of snap traps of Dionaea and Aldrovanda from a flypaper trap like Drosera regia, based on molecular data. The molecular and physiological data imply that Dionaea and Aldrovanda snap traps evolved from the flypaper traps of a common ancestor with Drosera. Pre-adaptations to the evolution of snap traps were identified in several species of Drosera, such as rapid leaf and tentacle movement. The model proposes that plant carnivory by snap trap evolved from the flypaper traps, driven by increasing prey size. Bigger prey provides greater nutritional value, but large insects can easily escape the sticky mucilage of flypaper traps; the evolution of snap traps would therefore prevent escape and kleptoparasitism (theft of prey captured by the plant before it can derive benefit from it), and would also permit a more complete digestion.[26][27]
In 2016, a study of the expression of genes in the plant's leaves as they captured and digested prey was published in the journal, Genome Research. The gene activation observed in the leaves of the plants gives support to the hypothesis that the carnivorous mechanisms present in the flytrap are a specially adapted version of mechanisms used by non-carnivorous plants to defend against herbivorous insects.[21][29] In many non-carnivorous plants, jasmonic acid serves as a signaling molecule for the activation of defense mechanisms, such as the production of hydrolases, which can destroy chitin and other molecular components of insect and microbial pests.[30] In the Venus flytrap, this same molecule has been found to be responsible for the activation of the plant's digestive glands. A few hours after the capture of prey, another set of genes is activated inside the glands, the same set of genes that is active in the roots of other plants, allowing them to absorb nutrients. The use of similar biological pathways in the traps as non-carnivorous plants use for other purposes indicates that somewhere in its evolutionary history, the Venus flytrap repurposed these genes for the purpose of carnivory.
Proposed evolutionary history
Carnivorous plants are generally herbs, and their traps the result of primary growth. They generally do not form readily fossilizable structures such as thick bark or wood. As a result, there is no fossil evidence of the steps that might link Dionaea and Aldrovanda, or either genus with their common ancestor, Drosera. Nevertheless, it is possible to infer an evolutionary history based on phylogenetic studies of both genera. Researchers have proposed a series of steps that would ultimately result in the complex snap-trap mechanism:[26][27]
Larger insects usually walk over the plant, instead of flying to it,[31] and are more likely to break free from sticky glands alone. Therefore, a plant with wider leaves, like Drosera falconeri,[26] must have adapted to move the trap and its stalks in directions that maximized its chance of capturing and retaining such prey—in this particular case, longitudinally. Once adequately "wrapped", escape would be more difficult.[31]
Evolutionary pressure then selected for plants with shorter response time, in a manner similar to Drosera burmannii or Drosera glanduligera. The faster the closing, the less reliant on the flypaper model the plant would be.
As the trap became more and more active, the energy required to "wrap" the prey increased. Plants that could somehow differentiate between actual insects and random detritus/rain droplets would have an advantage, thus explaining the specialization of inner tentacles into trigger hairs.
Ultimately, as the plant relied more on closing around the insect rather than gluing them to the leaf surface, the tentacles so evident in Drosera would lose their original function altogether, becoming the "teeth" and trigger hairs—an example of natural selection utilizing pre-existing structures for new functions.
Completing the transition, the plant eventually developed the depressed digestive glands found inside the trap, rather than using the dews in the stalks, further differentiating it from genus Drosera.
Habitat
Map of the original distribution of the Venus flytrap
The Venus flytrap is found in nitrogen- and phosphorus-poor environments, such as bogs and wet savannahs. Small in stature and slow-growing, the Venus flytrap tolerates fire well, and depends on periodic burning to suppress its competition.[32] Fire suppression threatens its future in the wild.[33] It survives in wet sandy and peaty soils. Although it has been successfully transplanted and grown in many locales around the world, it is native only to the coastal bogs of North and South Carolina in the United States, specifically within a 60-mile radius of Wilmington, North Carolina.[34] One such place is North Carolina's Green Swamp. There also appears to be a naturalized population of Venus flytraps in northern Florida as well as an introduced population in western Washington.[35][36] The nutritional poverty of the soil is the reason that the plant relies on such elaborate traps: insect prey provide the nitrogen for protein formation that the soil cannot. The Venus flytrap is not a tropical plant and can tolerate mild winters. In fact, Venus flytraps that do not go through a period of winter dormancy will weaken and die after a period of time.[37]
Cultivation
Dionaea muscipula 'Akai Ryu', Japanese for 'Red Dragon', in cultivation
Venus flytraps are popular as cultivated plants, but have a reputation for being difficult to grow.[8] Successfully growing these specialized plants requires recreating a close approximation to the plant's natural habitat.
Healthy Venus flytraps will produce scapes of white flowers in spring; however, many growers remove the flowering stems early (2–3 inches), as flowering consumes some of the plant's energy and thereby reduces the rate of trap production. If healthy plants are allowed to flower, successful pollination will result in seeds.
Plants can be propagated by seed, taking around four to five years to reach maturity. More commonly, they are propagated by clonal division in spring or summer. Venus flytraps can also be propagated in vitro using plant tissue culture.[38] Most Venus flytraps found for sale in nurseries garden centers have been produced using this method, as this is the most cost-effective way to propagate them on a large scale. Regardless of the propagation method used, the plants will live for 20 to 30 years if cultivated in the right conditions.[39]
Cultivars
Main article: List of Venus flytrap cultivars
Venus flytraps are by far the most commonly recognized and cultivated carnivorous plant, and they are frequently sold as houseplants. Various cultivars (cultivated varieties) have come into the market through tissue culture of selected genetic mutations, and these plants are raised in large quantities for commercial markets.
Conservation
The species is classified as "vulnerable" by the National Wildlife Federation.[40] In 2015, there were estimated to be fewer than 33,000 plants in the wild, all within 75 miles (121 km) of the city of Wilmington, North Carolina, and all on sites owned by The Nature Conservancy, the North Carolina state government, or the US military.[41]
In 2014, the state of North Carolina passed Senate Bill 734 which classifies the theft of naturally growing Venus flytraps in the state as a felony. Tougher sanctions and penalties for the theft were also enacted in December 1, 2014 in accordance with legislation.[42]
In alternative medicine
Venus flytrap extract is available on the market as an herbal remedy, sometimes as the prime ingredient of a patent medicine named "Carnivora". According to the American Cancer Society, these products are promoted in alternative medicine as a treatment for a variety of human ailments including HIV, Crohn's disease and skin cancer, but "available scientific evidence does not support the health claims made for Venus flytrap extract".[43]
See also
Carnivorous plants of North America
List of ineffective cancer treatments
References
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^ Jump up to: a b Böhm, J.; Scherzer, S.; Krol, E.; Kreuzer, I.; von Meyer, K.; Lorey, C.; Mueller, T. D.; Shabala, L.; Monte, I.; Solano, R.; Al-Rasheid, K. A. S.; Rennenberg, H.; Shabala, S.; Neher, E.; Hedrich, R. (2016). "The Venus Flytrap Dionaea muscipula Counts Prey-Induced Action Potentials to Induce Sodium Uptake". Current Biology. 26: 286–295. doi:10.1016/j.cub.2015.11.057. ISSN 0960-9822. PMC 4751343  . PMID 26804557.
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External links
Wikispecies has information related to: Dionaea muscipula
Wikimedia Commons has media related to Dionaea muscipula.
Images and movies of the Venus flytrap (Dionaea muscipula) at ARKive
Venus Flytrap Growing Guide and Distribution Map
The Carnivorous Plant FAQ: Flytraps
Venus flytrap origins uncovered - BBC
The CP Photo Finder: "Dionaea"
Dionaea muscipula - The Venus flytrap at Botanical Society of America
Criminal Podcast Episode Five: Dropping Like Flies
[hide]
v
t
e
Carnivorous and protocarnivorous plants
Carnivorous genera
(† extinct)
Aldrovanda
†Archaeamphora
Brocchinia
Byblis
Catopsis
Cephalotus
Darlingtonia
Dionaea
Drosera
†Droserapites
†Droserapollis
†Droseridites
Drosophyllum
†Fischeripollis
Genlisea
Heliamphora
Nepenthes
†Palaeoaldrovanda
Philcoxia
Pinguicula
Roridula
Sarracenia
†Saxonipollis
Triphyophyllum
Utricularia
Protocarnivorous genera
Aracamunia
Capsella
Colura
Dipsacus
Drymocallis
Geranium
Ibicella
Lathraea
Paepalanthus
Passiflora
Plumbago
Proboscidea
Stylidium
See also
International Carnivorous Plant Society
Insectivorous Plant Society
List of carnivorous plants
List of carnivorous plant periodicals
Pitcher plant
Taxon identifiers
Wd: Q155825
EoL: 584643
FNA: 220004158
GBIF: 5421410
GRIN: 405268
iNaturalist: 52666
IPNI: 275898-2
ITIS: 22008
IUCN: 39636
NCBI: 4362
Plant List: kew-62109
PLANTS: DIMU4
Species+: 21371
Tropicos: 11400032
ARKive: venus-flytrap/dionaea-muscipula
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Mobile network operators of India
From Wikipedia, the free encyclopedia
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This article needs to be updated. Please update this article to reflect recent events or newly available information. (August 2017)
This is a list of mobile network operators of India as on 31 July 2017.[1]

Rank
Operator's Name
Technology
Subscribers[1]
(in millions)
Market Share[1]
Ownership
1 VODAFONE/IDEA * Vodafone India - GSM-900/1800 (EDGE)
2100 MHz UMTS, HSPA+, EVDO
1800 MHz LTE
* Idea Cellular - GSM-900/1800 (EDGE)
2100 MHz UMTS, HSPA+
LTE Band 3 (1800 MHz)[2] * Vodafone India - 210.54
* Idea Cellular - 193.96 * Vodafone India - 17.74%
* Idea Cellular - 16.34% Vodafone Group (45.1%)
Aditya Birla Group (26%)
Axiata Group Berhad
Providence Equity
2 AIRTEL GSM-900/1800 (EDGE)
2100 MHz UMTS, HSPA+
LTE Band 3 (1800 MHz),
LTE Band 40 (2300 MHz),
Voice over LTE /VoLTE (Testing Phase) LTE Advanced
281.25 23.70% Bharti Enterprises (64%)
SingTel (36%)
3 Jio LTE Band 5 (850 MHz),
LTE Band 3 (1800 MHz),
LTE Band 40 (2300 MHz),
Voice over LTE / VoLTE
LTE Advance
138.60 10.83% Reliance Industries
4 BSNL Mobile GSM-900/1800 (EDGE)
2100 MHz UMTS, HSPA+
CdmaOne, EVDO
WiMAX WiFi 104.55 8.81% Government-owned
5 Aircel GSM-900/1800 (EDGE)
2100 MHz UMTS, HSPA+
2300 MHz TD-LTE 89.93 7.58% Maxis Communications (74%)
Sindya Securities and Investments (26%)
6 Reliance Communications GSM-900/1800 (EDGE)
2100 MHz UMTS, HSPA+
WiMAX 81.27 6.85% Reliance ADAG
7 Telenor India GSM-1800 MHz (EDGE)
2300 MHz TD-LTE 47.06 3.97% Telenor Group
8 Tata Teleservices GSM-1800 (EDGE)
2100 MHz UMTS, HSPA+
CDMA, EVDO
WiMAX 42.09 3.55% Tata Teleservices (100%)
9 MTS India CDMA2000, EVDO 3.92 0.33% Sistema (56.68%)
Shyam Group (23.98)
Government of Russia (17.14%)
10 MTNL GSM-900 MHz (EDGE)
2100 MHz UMTS, HSPA 3.62 0.30% State-owned
Note
Airtel is acquiring Telenor[3] and Tata Teleservices.[4]
Vodafone and Idea have still not merged [5]
Reliance Communications and MTS are merging. [6]
A merger of state-owned telecom companies BSNL and MTNL is being mulled over by the Indian Government.[7]
See also[edit]
Telecommunications in India
List of telecom companies in India
List of mobile network operators
List of mobile network operators of the Asia Pacific region
References[edit]
^ Jump up to: a b c "Press Release on Telecom Subscription Data". Telecom Regulatory Authority of India.
Jump up ^ "Idea covers 10 telecom circles with its world class 4G LTE services". Idea Cellular.
Jump up ^ "Bharti Airtel to acquire Telenor India - The Economic Times". The Economic Times. Retrieved 2017-03-21.
Jump up ^ "Tata Teleservices to sell its wireless mobile business to Bharti Airtel". The Economic Times. October 13, 2017.
Jump up ^ "Vodafone, Idea Agree on Merger to Create Indian Wireless Leader". Bloomberg.com. 2017-03-20. Retrieved 2017-03-21.
Jump up ^ SH, Salman (2016-10-03). "Update: RCOM and SSTL merger gets approval from Rajasthan HC". MediaNama. Retrieved 2017-03-21.
Jump up ^ "MTNL, BSNL revival on cards? This is what Narendra Modi government is planning". Financial Express. October 11, 2017.
[show]
v
t
e
Telecommunication companies of India
Mobile network operators
Aircel
Airtel
BSNL Mobile
Idea
Jio
MTNL
Reliance
T24 Mobile
Tata DoCoMo
Telenor
Vodafone
Defunct mobile operators
Loop Mobile
MTS
Orange
S Tel
Spice Telecom
Videocon
Virgin Mobile
Internet service providers
ACT Fibernet
Airtel Broadband
APSFL
BBNL
BSNL
Hathway
Jio
MTNL
Railtel
Reliance
Sify
Spectranet
Tata Teleservices
Tikona Broadband
DTH operators
ABS Free Dish
ACT Digital
Airtel digital TV
DD Free Dish
Dish TV
Reliance Digital TV
Sun Direct
Tata Sky
Videocon d2h
Zing Digital
Retrieved from "https://en.wikipedia.org/w/index.php?title=Mobile_network_operators_of_India&oldid=807215047"
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Mobile phone companies of India
Internet service providers of India
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