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Animal communication
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Great egret (Ardea alba) in a courtship display communicating the desire to find a mate
Animal communication is the transfer of information from one or a group of animals (sender or senders) to one or more other animals (receiver or receivers) that affects the current or future behavior of the receivers. Information may be sent intentionally, as in a courtship display, or unintentionally, as in the transfer of scent from predator to prey. Information may be transferred to an "audience" of several receivers. Animal communication is a rapidly growing area of study in disciplines including animal behavior, sociology, neurology and animal cognition. Many aspects of animal behavior, such as symbolic name use, emotional expression, learning and sexual behavior, are being understood in new ways.
When the information from the sender changes the behavior of a receiver, the information is referred to as a "signal". Signalling theory predicts that for a signal to be maintained in the population, both the sender and receiver should usually receive some benefit from the interaction. Signal production by senders and the perception and subsequent response of receivers are thought to coevolve.[1] Signals often involve multiple mechanisms, e.g. both visual and auditory, and for a signal to be understood the coordinated behaviour of both sender and receiver require careful study.
Contents
1 Modes
1.1 Visual
1.2 Auditory
1.3 Olfactory
1.4 Electric
1.5 Touch
1.6 Seismic
1.7 Thermal
2 Autocommunication
3 Functions
4 Interpretation of animal behaviour
5 Interspecific communication
5.1 Prey to predator
5.2 Predator to prey
5.3 Human/animal
6 Other aspects
6.1 Evolution
6.2 Cognitive aspects
6.3 Human behaviour
6.4 Linguistics
7 Errors in communication
8 See also
9 References
10 External links
Modes[edit]
A lamb investigates a rabbit, an example of interspecific communication using body posture and olfaction.
Visual[edit]
For information on the perception of visual signals, see Visual perception.
Gestures: The best known form of communication involves the display of distinctive body parts, or distinctive bodily movements; often these occur in combination, so a movement acts to reveal or emphasize a body part. A notable example is the presentation of a parent herring gull’s bill to its chick as a signal for feeding. Like many gulls, the herring gull has a brightly coloured bill, yellow with a red spot on the lower mandible near the tip. When the parent returns to the nest with food, it stands over its chick and taps the bill on the ground; this elicits a begging response from a hungry chick (pecking at the red spot), which stimulates the parent to regurgitate food. The complete signal therefore involves a distinctive morphological feature (body part), the red-spotted bill, and a distinctive movement (tapping towards the ground) which makes the red spot highly visible to the chick. While all primates use some form of gesture,[2] Frans de Waal concluded that apes and humans are unique in that only they use intentional gestures to communicate. He tested the hypothesis that gestures evolve into language by studying the gestures of bonobos and chimps.
A dog's visual tooth-baring threat display, accompanied by an auditory signal, growling
Facial expression: Facial gestures play an important role in animal communication. Often a facial gesture is a signal of emotion. Dogs, for example, express anger through snarling and showing their teeth. In alarm their ears perk up, in fear the ears flatten while the dogs expose their teeth slightly and squint their eyes. Jeffrey Mogil studied the facial expressions of mice during increments of increasing pain; there were five recognizable facial expressions; orbital tightening, nose and cheek bulge, and changes in ear and whisker carriage.[3]
Gaze following: Social animals coordinate their communication by monitoring of each other's head and eye orientation. Such behaviour has long been recognized as an important component of communication during human development, and gaze-following has recently received much attention in animals. Studies have been conducted on apes, monkeys, dogs, birds, and tortoises, and have focused on two different tasks: "follow[ing] another’s gaze into distant space" and "follow[ing] another’s gaze geometrically around a visual barrier e.g. by repositioning themselves to follow a gaze cue when faced with a barrier blocking their view". The first ability has been found among a broad range of animals, while the second has been demonstrated only for apes, dogs, wolves, and corvids (ravens); attempts to demonstrate this "geometric gaze following" in marmoset and ibis gave negative results. Researchers do not yet have a clear picture of the cognitive basis of gaze following, but developmental evidence indicates that "simple" gaze following and "geometric" gaze following probably rely on different cognitive mechanisms.[4]
Color change: Color change can be separated into changes that occur during growth and development, and those triggered by mood, social context, or abiotic factors such as temperature. The latter are seen in many taxa. Some cephalopods, such as the octopus and the cuttlefish, have specialized skin cells (chromatophores) that can change the apparent colour, opacity, and reflectiveness of their skin.[5] In addition to their use for camouflage, rapid changes in skin color are used while hunting and in courtship rituals.[6] Cuttlefish may display two entirely different signals simultaneously from opposite sides of their body. When a male cuttlefish courts a female in the presence of other males, he displays a male pattern facing the female and a female pattern facing away, to deceive other males.[7] Some color signals occur in cycles. For example, when a female olive baboon begins to ovulate, her anogenital area swells and turns a bright red/pink. This signals to males that she is ready to mate.[8]
Bioluminescent communication: Communication by the production of light occurs commonly in vertebrates and invertebrates in the oceans, particularly at depths (e.g. angler fish). Two well known forms of land bioluminescence occur in fireflies and glow worms. Other insects, insect larvae, annelids, arachnids and even species of fungi possess bioluminescent abilities. Some bioluminescent animals produce the light themselves whereas others have a symbiotic relationship with bioluminescent bacteria. (See also: List of bioluminescent organisms)
Auditory[edit]
Further information: bird song
Bird calls can serve as alarms or keep members of a flock in contact, while the longer and more complex bird songs are associated with courtship and mating.[9]
Crows flocking
A flock of American crows.
Problems playing this file? See media help.
Humpback Whale singing at southern ocean feeding grounds.
Many animals communicate through vocalization. Vocal communication serves many purposes, including mating rituals, warning calls, conveying location of food sources, and social learning. In a number of species, males perform calls during mating rituals as a form of competition against other males and to signal females. Examples include frogs, hammer-headed bats, red deer, humpback whales, elephant seals, and songbirds.[10] Other instances of vocal communication include the alarm calls of the Campbell monkey,[11] the territorial calls of gibbons, and the use of frequency in greater spear-nosed bats to distinguish between groups.[12] The vervet monkey gives a distinct alarm call for each of its four different predators, and the reactions of other monkeys vary appropriately according to the call. For example, if an alarm call signals a python, the monkeys climb into the trees, whereas the "eagle" alarm causes monkeys to seek a hiding place on the ground. Prairie dogs also use complex calls that signal predator differences. According to Con Slobodchikoff and others, prairie dog calls communicate the type, size, and speed of an approaching predator.[13][14][15][16] Whale vocalizations have been found to have different dialects based on region.[17]
Not all animals use vocalization as a means of auditory communication. Many arthropods rub specialized body parts together to produce sound. This is known as stridulation. Crickets and grasshoppers are well known for this, but many others use stridulation as well, including crustaceans, spiders, scorpions, wasps, ants, beetles, butterflies, moths, millipedes, and centipedes. Another means of auditory communication is the vibration of swim bladders in bony fish. The structure of swim bladders and the attached sonic muscles varies greatly across bony fish families, resulting in a wide variety of sounds. Striking body parts together can also produce auditory signals. A well-known example of this is the tail tip vibration of rattlesnakes as a warning signal. Other examples include bill clacking in birds, wing clapping in manakin courtship displays, and chest beating in gorillas.[18]
Olfactory[edit]
Main article: Olfaction
Despite being the oldest method of communication, chemical communication is one of the least understood forms due in part to the sheer abundance of chemicals in our environment and the difficulty of detecting and measuring all the chemicals in a sample.[18] The ability to detect chemicals in the environment serves many functions, a crucial one being the detection of food, a function that first arose in single-celled organisms (bacteria) living in the oceans during the early days of life on Earth.[18] As this function evolved, organisms began to differentiate between chemicals compounds emanating from resources, conspecifics (same species; i.e., mates and kin), and heterospecifics (different species; i.e., competitors and predators).[18] For instance, a small minnow species may do well to avoid habitat with a detectable concentration of chemical cue associated with a predator species such as northern pike.[19] Minnows with the ability to perceive the presence of predators before they are close enough to be seen and then respond with adaptive behaviour (such as hiding) are more likely to survive and reproduce.[citation needed] Scent marking and scent rubbing are common forms of olfactory communication in mammals.[20][21]
Electric[edit]
Main article: Electrocommunication
Electrocommunication is a rare form of communication in animals. It is seen primarily in aquatic animals, though some land mammals, notably the platypus and echidnas, sense electric fields that might be used for communication.[22]
Weakly electric fishes provide an example of electrocommunication, together with electrolocation. These fish use an electric organ to generate an electric field, which is detected by electroreceptors. Differences in the waveform and frequency of changes in the field convey information on species, sex, and identity. These electric signals can be generated in response to hormones, circadian rhythms, and interactions with other fish. Some predators, such as sharks and rays, are able to eavesdrop on these electrogenic fish through passive electroreception.
Touch[edit]
For more on the mechanism for touch, see Somatosensory system and Mechanoreceptors
Touch is a key factor in many social interactions. Here are some examples:
Fighting: In a fight, touch may be used to challenge an opponent and to coordinate movements during the fight. It may also be used by the loser to indicate submission.
Mating: Mammals often initiate mating by grooming, stroking or rubbing against each other. This provides the opportunity to apply chemical signals and to assess those excreted by the potential mate. Touch may also announce the intention of the male to mount the female, as when a male kangaroo grabs the tail of a female. During mating, touch stimuli are important for pair positioning, coordination and genital stimulation.
Social integration: Touch is widely used for social integration, a use that is typified by the social grooming of one animal by another. Social grooming has several functions; it removes parasites and debris from the groomed animal, it reaffirms the social bond or hierarchical relationship between the animals, and it gives the groomer an opportunity to examine olfactory cues on the groomed individual, perhaps adding additional ones. This behaviour has been observed in social insects, birds and mammals.
Foraging: Some ant species recruit fellow workers to new food finds by first tapping them with their antennae and forelegs, then leading them to the food source while keeping physical contact. Another example of this is the waggle dance of honey bees.[23]
Huddling: Prolonged physical contact or huddling also serves social integration. Huddling promotes heat exchange, together with the transfer of olfactory or tactile information. Some organisms live in constant contact in a colony, for example colonial corals. When individuals are linked tightly in this way an entire colony can react on the aversive or alarm movements made by only a few individuals. In several herbivorous insect nymphs and larvae, aggregations where there is prolonged contact play a major role in group coordination. These aggregations may take the form of a procession or a rosette.
Seismic[edit]
Main article: Seismic communication
Seismic communication is the exchange of information using self-generated vibrational signals transmitted via a substrate such as the soil, water, spider webs, plant stems, or a blade of grass. This form of communication has several advantages, for example it can be sent regardless of light and noise levels, and it usually has a short range and short persistence, which may reduce the danger of detection by predators. The use of seismic communication is found in many taxa, including frogs, kangaroo rats, mole rats, bees, nematode worms, and others. Tetrapods usually make seismic waves by drumming on the ground with a body part, a signal that is sensed by the sacculus of the receiver. The sacculus is an organ in the inner ear containing a membranous sac that is used for balance, but can also detect seismic waves in animals that use this form of communication. Vibrations may be combined with other sorts of communication.[24]
Thermal[edit]
Main article: Thermoreception
A python (top) and rattlesnake illustrating the positions of the pit organs. Red arrows indicate the pit organs whereas black arrows indicate the nostril.
A number of different snakes have the ability to sense infrared (IR) thermal radiation, which allows these reptiles to derive thermal images from the radiant heat emitted by predators or prey at wavelengths between 5 and 30 μm. The accuracy of this sense is such that a blind rattlesnake can target its strike to the vulnerable body parts of a prey animal.[25] It was previously thought that the pit organs evolved primarily as prey detectors, but it is now believed that they may also be used to control body temperature.[26]
The facial pits enabling thermoregulation underwent parallel evolution in pitvipers and some boas and pythons, having evolved once in pitvipers and multiple times in boas and pythons.[27] The electrophysiology of the structure is similar between lineages, but it differs in gross structure anatomy. Most superficially, pitvipers possess one large pit organ on either side of the head, between the eye and the nostril (loreal pit), while boas and pythons have three or more comparatively smaller pits lining the upper and sometimes the lower lip, in or between the scales. Those of the pitvipers are the more advanced, having a suspended sensory membrane as opposed to a simple pit structure. Within the family Viperidae, the pit organ is seen only in the subfamily Crotalinae: the pitvipers. Despite the detection of IR radiation, the pits’ IR mechanism is dissimilar to photoreceptors; while photoreceptors detect light via photochemical reactions, the protein in the facial pits of snakes is a temperature sensitive ion channel. It senses infrared signals through a mechanism involving warming of the pit organ, rather than chemical reaction to light.[28] This is consistent with the thin pit membrane, which allows incoming IR radiation to quickly and precisely warm a given ion channel and trigger a nerve impulse, as well as vascularize the pit membrane to rapidly cool the ion channel back to its original “resting” or “inactive” temperature.[28]
Common vampire bats (Desmodus rotundus) have specialized IR sensors in their nose-leaf.[29] Vampire bats are the only mammals that feed exclusively on blood. The IR sense enables Desmodus to localize homeothermic animals such as cattle and horses within a range of about 10 to 15 cm. This infrared perception may be used in detecting regions of maximal blood flow on targeted prey.
Autocommunication[edit]
Autocommunication is a type of communication in which the sender and receiver are the same individual. The sender emits a signal that is altered by the environment and eventually is received by the same individual. The altered signal provides information that can indicate food, predators or conspecifics. Because the sender and receiver are the same animal, selection pressure maximizes signal efficacy, i.e. the degree to which an emitted signal is correctly identified by a receiver despite propagation distortion and noise. There are two types of autocommunication. The first is active electrolocation found in the electric fish Gymnotiformes (knifefishes) and Mormyridae (elephantfish) and also in the platypus (Ornithorhynchus anatinus). The second type of autocommunication is echolocation, found in bats and toothed whales.
Functions[edit]
There are many functions of animal communication. However, some have been studied in more detail than others. This includes:
Communication during contests: Animal communication plays a vital role in determining the winner of contest over a resource. Many species have distinct signals that signal aggression or willingness to attack or signals to convey retreat during competitions over food, territories, or mates.
Two Red Deer roaring most likely to establish dominance during a rut. However, males also use loud roaring to keep track of harems of females.
Mating rituals: Animals produce signals to attract the attention of a possible mate or to solidify pair bonds. These signals frequently involve the display of body parts or postures. For example, a gazelle will assume characteristic poses to initiate mating. Mating signals can also include the use of olfactory signals or mating calls unique to a species. Animals that form lasting pair bonds often have symmetrical displays that they make to each other. Famous examples are the mutual presentation of reeds by great crested grebes studied by Julian Huxley, the triumph displays shown by many species of geese and penguins on their nest sites, and the spectacular courtship displays by birds of paradise.
Ownership/territorial: Signals used to claim or defend a territory, food, or a mate.
Food-related signals: Many animals make "food calls" to attract a mate, offspring, or other members of a social group to a food source. Perhaps the most elaborate food-related signal is the Waggle dance of honeybees studied by Karl von Frisch. One well-known example of begging of offspring in a clutch or litter is altricial songbirds. Young ravens signal will signal to older ravens when they encounter new or untested food. Rhesus macaques will send food calls to inform other monkeys of a food source to avoid punishment. Pheromones are released by many social insects to lead the other members of the society to the food source. For example, ants leave a pheromone trail on the ground that can be followed by other ants to lead them to the food source.
Alarm calls: Alarm calls communicate the threat of a predator. This allows all members of a social group (and sometimes other species) to respond accordingly. This may include running for cover, becoming immobile, or gathering into a group to reduce the risk of attack. Alarm signals are not always vocalizations. Crushed ants will release an alarm pheromone to attract more ants and send them into an attack state.
Meta-communication: Signals that will modify the meaning of subsequent signals. One example is the 'play face' in dogs which signals that a subsequent aggressive signal is part of a play fight rather than a serious aggressive episode.
Interpretation of animal behaviour[edit]
As described above, many animal gestures, postures, and sounds, convey meaning to nearby animals. These signals are often easier to describe than to interpret. It is tempting, especially with domesticated animals and apes, to anthropomorphize, that is, to interpret animal actions in human terms, but this can be quite misleading; for example, an ape's "smile" is often a sign of aggression. Also, the same gesture may have different meanings depending on context within which it occurs. For example, a domestic dog's tail wag and posture may be used in different ways to convey many meanings as illustrated in Charles Darwin's The Expression of the Emotions in Man and Animals published in 1872. Some of Darwin's illustrations are reproduced here.
Examples of tail position indicating different emotions in dogs
"Small dog watching a cat on a table"
"Dog approaching another dog with hostile intentions"
"Dog in a humble and affectionate frame of mind"
"Half-bred shepherd dog"
"Dog caressing his master"
Interspecific communication[edit]
Main article: Interspecies communication
Much animal communication is intraspecific, that is, it occurs between members of the same species. As for interspecific communication, that between predator and prey is of particular interest.
Prey to predator[edit]
If a prey animal moves, makes a noise or vibrations, or emits a smell in such a way that a predator can detect it, this is consistent with the definition of "communication" given above. This type of communication is known as interceptive eavesdropping if a predator intercepts a message intended for conspecifics.
There are however, some actions of prey species are clearly directed to actual or potential predators. A good example is warning coloration: species such as wasps that are capable of harming potential predators are often brightly coloured, and this modifies the behaviour of the predator, who either instinctively or as the result of experience will avoid attacking such an animal. Some forms of mimicry fall in the same category: for example hoverflies are coloured in the same way as wasps, and although they are unable to sting, the strong avoidance of wasps by predators gives the hoverfly some protection. There are also behavioural changes that act in a similar way to warning colouration. For example, canines such as wolves and coyotes may adopt an aggressive posture, such as growling with their teeth bared, to indicate they will fight if necessary, and rattlesnakes use their well-known rattle to warn potential predators of their venomous bite. Sometimes, a behavioural change and warning colouration will be combined, as in certain species of amphibians which have most of their body coloured to blend with their surroundings, except for a brightly coloured belly. When confronted with a potential threat, they show their belly, indicating that they are poisonous in some way.
Another example of prey to predator communication is the pursuit-deterrent signal. Pursuit-deterrent signals occur when prey indicates to a predator that pursuit would be unprofitable because the signaler is prepared to escape. Pursuit-deterrent signals provide a benefit to both the signaler and receiver; they prevent the sender from wasting time and energy fleeing, and they prevent the receiver from investing in a costly pursuit that is unlikely to result in capture. Such signals can advertise prey’s ability to escape, and reflect phenotypic condition (quality advertisement), or can advertise that the prey has detected the predator (perception advertisement). Pursuit-deterrent signals have been reported for a wide variety of taxa, including fish (Godin and Davis, 1995), lizards (Cooper et al., 2004), ungulates (Caro, 1995), rabbits (Holley 1993), primates (Zuberbuhler et al. 1997), rodents (Shelley and Blumstein 2005, Clark, 2005), and birds (Alvarez, 1993, Murphy, 2006, 2007). A familiar example of quality advertisement pursuit-deterrent signal is stotting (sometimes called pronking), a pronounced combination of stiff-legged running while simultaneously jumping shown by some antelopes such as Thomson's gazelle in the presence of a predator. At least 11 hypotheses for stotting have been proposed. A leading theory today is that it alerts predators that the element of surprise has been lost. Predators like cheetahs rely on surprise attacks, proven by the fact that chases are rarely successful when antelope stot. Predators do not waste energy on a chase that will likely be unsuccessful (optimal foraging behaviour). Quality advertisement can be communicated by modes other than visual. The banner-tailed kangaroo rat produces several complex foot-drumming patterns in a number of different contexts, one of which is when it encounters a snake. The foot-drumming may alert nearby offspring but most likely conveys vibrations through the ground that the rat is too alert for a successful attack, thus preventing the snake's predatory pursuit.[30]
Predator to prey[edit]
The humpback anglerfish angles for small fish by deceptively dangling a bioluminescent lure in front of its jaws.
Further information: Deception in animals
Typically, predators attempt to reduce communication to prey as this will generally reduce the effectiveness of their hunting. However, some forms of predator to prey communication occur in ways that change the behaviour of the prey and make their capture easier, i.e. deception by the predator. A well-known example is the angler fish, an ambush predator which waits for its prey to come to it. It has a fleshy bioluminescent growth protruding from its forehead which it dangles in front of its jaws. Smaller fish attempt to take the lure, placing themselves in a better position for the angler fish to catch them. Another example of deceptive communication is observed in the genus of jumping spiders (Myrmarachne). These spiders are commonly referred to as “antmimicking spiders” because of the way they wave their front legs in the air to simulate antennae.
Human/animal[edit]
Main article: Human–animal communication
Various ways in which humans interpret the behaviour of animals, or give commands to them, are consistent with the definition of interspecies communication. Skillful interpretation of animal communications may be critical to the welfare of animals that are being cared for or trained by humans. For example, behaviours indicating pain need to be recognized. Indeed, the survival of both the animal and its human caretaker may be at stake if, for example, a human fails to recognize a signal for imminent attack. It is also important to take into account that non-human animal species may interpret the signals of humans differently than humans themselves. For instance, a pointing command refers to a location rather than an object in dogs[31].
Since the late 1990s, one scientist, Sean Senechal, has been developing, studying, and using the learned visible, expressive language in dogs and horses. By teaching these animals a gestural (human made) American Sign Language-like language, the animals have been found to use the new signs on their own to get what they need.[32] The recent experiments on animal language are perhaps the most sophisticated attempt yet to establish human/animal communication, though their relation to natural animal communication is uncertain.
Other aspects[edit]
Evolution[edit]
Further information: Signalling theory
The importance of communication is evident from the highly elaborate morphology, behaviour and physiology that some animals have evolved to facilitate this. These include some of the most striking structures in the animal kingdom, such as the peacock's tail, the antlers of a stag and the frill of the frill-necked lizard, but also include even the modest red spot on a European herring gull's bill. Highly elaborate behaviours have evolved for communication such as the dancing of cranes, the pattern changes of cuttlefish, and the gathering and arranging of materials by bowerbirds. Other evidence for the importance of communication in animals is the prioritisation of physiological features to this function, for example, birdsong appears to have brain structures entirely devoted to its production. All these adaptations require evolutionary explanation.
There are two aspects to the required explanation:
identifying a route by which an animal that lacked the relevant feature or behaviour could acquire it;
identifying the selective pressure that makes it adaptive for animals to develop structures that facilitate communication, emit communications, and respond to them.
Significant contributions to the first of these problems were made by Konrad Lorenz and other early ethologists. By comparing related species within groups, they showed that movements and body parts that in the primitive forms had no communicative function could be "captured" in a context where communication would be functional for one or both partners, and could evolve into a more elaborate, specialised form. For example, Desmond Morris showed in a study of grass finches[citation needed] that a beak-wiping response occurred in a range of species, serving a preening function, but that in some species this had been elaborated into a courtship signal.
The second problem has been more controversial. The early ethologists assumed that communication occurred for the good of the species as a whole, but this would require a process of group selection which is believed to be mathematically impossible in the evolution of sexually reproducing animals. Altruism towards an unrelated group is not widely accepted in the scientific community, but rather can be seen as reciprocal altruism, expecting the same behaviour from others, a benefit of living in a group. Sociobiologists argued that behaviours that benefited a whole group of animals might emerge as a result of selection pressures acting solely on the individual. A gene-centered view of evolution proposes that behaviours that enabled a gene to become wider established within a population would become positively selected for, even if their effect on individuals or the species as a whole was detrimental;[33]
The apparently excessive eye-spot signalling by the male peacock tail may be runaway selection
In the case of communication, an important discussion by John Krebs and Richard Dawkins established hypotheses for the evolution of such apparently altruistic or mutualistic communications as alarm calls and courtship signals to emerge under individual selection. This led to the realization that communication might not always be "honest" (indeed, there are some obvious examples where it is not, as in mimicry). The possibility of evolutionarily stable dishonest communication has been the subject of much controversy, with Amotz Zahavi in particular arguing that it cannot exist in the long term. Sociobiologists have also been concerned with the evolution of apparently excessive signaling structures such as the peacock's tail; it is widely thought that these can only emerge as a result of sexual selection, which can create a positive feedback process that leads to the rapid exaggeration of a characteristic that confers an advantage in a competitive mate-selection situation.
One theory to explain the evolution of traits like a peacock's tail is 'runaway selection'. This requires two traits—a trait that exists, like the bright tail, and a prexisting bias in the female to select for that trait. Females prefer the more elaborate tails, and thus those males are able to mate successfully. Exploiting the psychology of the female, a positive feedback loop is enacted and the tail becomes bigger and brighter. Eventually, the evolution will level off because the survival costs to the male do not allow for the trait to be elaborated any further. Two theories exist to explain runaway selection. The first is the good genes hypothesis. This theory states that an elaborate display is an honest signal of fitness and truly is a better mate. The second is the handicap hypothesis. This explains that the peacock's tail is a handicap, requiring energy to keep and makes it more visible to predators. Thus, the signal is costly to maintain, and remains an honest indicator of the signaler's condition. Another assumption is that the signal is more costly for low quality males to produce than for higher quality males to produce. This is simply because the higher quality males have more energy reserves available to allocate to costly signaling.[1]
Cognitive aspects[edit]
Ethologists and sociobiologists have characteristically analysed animal communication in terms of more or less automatic responses to stimuli, without raising the question of whether the animals concerned understand the meaning of the signals they emit and receive. That is a key question in animal cognition. There are some signalling systems that seem to demand a more advanced understanding. A much discussed example is the use of alarm calls by vervet monkeys. Robert Seyfarth and Dorothy Cheney showed that these animals emit different alarm calls in the presence of different predators (leopards, eagles, and snakes), and the monkeys that hear the calls respond appropriately—but that this ability develops over time, and also takes into account the experience of the individual emitting the call. Metacommunication, discussed above, also seems to require a more sophisticated cognitive process.
It has been reported [34] that bottlenose dolphins can recognize identity information from whistles even when otherwise stripped of the characteristics of the whistle; making dolphins the only animals other than humans that have been shown to transmit identity information independent of the caller’s voice or location. The paper concludes that:
The fact that signature whistle shape carries identity information independent from voice features presents the possibility to use these whistles as referential signals, either addressing individuals or referring to them, similar to the use of names in humans. Given the cognitive abilities of bottlenose dolphins, their vocal learning and copying skills, and their fission–fusion social structure, this possibility is an intriguing one that demands further investigation.
—  V. M. Janik, et al. [34]
Human behaviour[edit]
Another controversial issue is the extent to which human behaviours resemble animal communication, or whether all such communication has disappeared as a result of our linguistic capacity. Some of our bodily features—eyebrows, beards and moustaches, deep adult male voices, perhaps female breasts—strongly resemble adaptations to producing signals. Ethologists such as Irenäus Eibl-Eibesfeldt have argued that facial gestures such as smiling, grimacing, and the eyebrow flash on greeting are universal human communicative signals that can be related to corresponding signals in other primates. Given how recently spoken language has emerged, it is very likely that human body language does include some more or less involuntary responses that have a similar origin to the communication we have.
Humans also often seek to mimic animals' communicative signals in order to interact with them. For example, cats have a mild affiliative response of slowly closing their eyes; humans often mimic this signal towards a pet cat to establish a tolerant relationship. Stroking, petting and rubbing pet animals are all actions that probably work through their natural patterns of interspecific communication.
Dogs have shown an ability to understand human communication. In object choice tasks, dogs utilize human communicative gestures such as pointing and direction of gaze in order to locate hidden food and toys.[35] It has also been shown that dogs exhibit a left gaze bias when looking at human faces, indicating that they are capable of reading human emotions.[36] Dogs do not make use of direction of gaze or exhibit left gaze bias with other dogs.
A new approach in the 21st century in the field of animal communication uses applied behavioural analysis, specifically functional communication training. This form of training previously has been used in schools and clinics with humans with special needs, such as children with autism, to help them develop language. Sean Senechal at the AnimalSign Center has been using an approach similar to functional communication training with domesticated animals, such as dogs since 2004 and horses since 2000, with encouraging results and benefits to the animals and people. Functional communication training for animals, Senechal calls "AnimalSign Language". This includes teaching communication through gestures (like simplified American sign language), Picture Exchange Communication System, tapping, and vocalisation. The process for animals includes simplified and modified techniques.[citation needed]
Linguistics[edit]
External video
Do animals have language? - Michele Bishop, TED Ed, 4:54, September 10, 2015[37]
Main article: Animal language
For linguistics, the interest of animal communication systems lies in their similarities to and differences from human language:
Human languages are characterized for having a double articulation (in the characterization of French linguist André Martinet). It means that complex linguistic expressions can be broken down in meaningful elements (such as morphemes and words), which in turn are composed of smallest phonetic elements that affect meaning, called phonemes. Animal signals, however, do not exhibit this dual structure.
In general, animal utterances are responses to external stimuli, and do not refer to matters removed in time and space. Matters of relevance at a distance, such as distant food sources, tend to be indicated to other individuals by body language instead, for example wolf activity before a hunt, or the information conveyed in honeybee dance language.It is therefore unclear to what extent utterances are automatic responses and to what extent deliberate intent plays a part.
In contrast to human language, animal communication systems are usually not able to express conceptual generalizations. (Cetaceans and some primates may be notable exceptions).
Human languages combine elements to produce new messages (a property known as creativity). One factor in this is that much human language growth is based upon conceptual ideas and hypothetical structures, both being far greater capabilities in humans than animals. This appears far less common in animal communication systems, although current research into animal culture is still an ongoing process with many new discoveries.
A recent and interesting area of development is the discovery that the use of syntax in language, and the ability to produce "sentences", is not limited to humans either. The first good evidence of syntax in non-humans, reported[38] in 2006, is from the greater spot-nosed monkey (Cercopithecus nictitans) of Nigeria. This is the first evidence that some animals can take discrete units of communication, and build them up into a sequence which then carries a different meaning from the individual "words":
The greater spot-nosed monkeys have two main alarm sounds. A sound known onomatopoeiacally as the "pyow" warns against a lurking leopard, and a coughing sound that scientists call a "hack" is used when an eagle is flying nearby.
"Observationally and experimentally we have demonstrated that this sequence [of up to three 'pyows' followed by up to four 'hacks'] serves to elicit group movement... the 'pyow-hack' sequence means something like 'let's go!' [a command telling others to move]... The implications are that primates at least may be able to ignore the usual relationship between an individual alarm call, and the meaning it might convey under certain circumstances... To our knowledge this is the first good evidence of a syntax-like natural communication system in a non-human species."
Similar results have also recently been reported in the Campbell's mona monkey.[39]
Errors in communication[edit]
There becomes possibility for error within communication between animals when certain circumstances apply. [40]These circumstances could include distance between the two communicating subjects, as well as the complexity of the signal that is being communicated to the "listener" of the situation. It may not always be clear to the "listener" where the location of the communication is coming from, as the "singer" can sometimes deceive them and create more error.[citation needed]
See also[edit]
Animals portal
Animal consciousness
Anthrozoology (human–animal studies)
Biocommunication
Biosemiotics
Body language
Dear enemy effect and Nasty neighbour effect
Deception in animals
Degeneracy (biology)
Emotion in animals
Forms of activity and interpersonal relations
Human–animal communication
International Society for Biosemiotic Studies
Origin of language
Origin of speech
Sir Philip Sidney game
Talking animal
Zoomusicology
Zoosemiotics
References[edit]
^ Jump up to: a b Maynard-Smith and Harper, 2003
Jump up ^ de Waal
Jump up ^ Langford, D.J. et al., (2010). Coding of facial expressions of pain in the laboratory mouse. Nature Methods, May 9th. pp. 1-3. doi:10.1038/nmeth.1455
Jump up ^ Range, F.; Virányi, Z. (2011). "Development of gaze following abilities in wolves (Canis Lupus)". PLoS ONE. 6 (2): e16888. Bibcode:2011PLoSO...616888R. doi:10.1371/journal.pone.0016888. PMC 3044139. PMID 21373192.
Jump up ^ Cloney, R.A.; Florey, E. (1968). "Ultrastructure of cephalopod chromatophore organs". Z. Zellforsch Mikrosk. Anat. 89: 250–280. doi:10.1007/bf00347297. PMID 5700268.
Jump up ^ Hanlon, R.T.; Messenger, J.B. (1996). Cephalopod Behaviour. Cambridge University Press. p. 121. ISBN 0-521-64583-2.
Jump up ^ Williams, Sarah (2012). "Two-faced fish tricks competitors". Science Now. Archived from the original on March 8, 2013. Retrieved March 16, 2013.
Jump up ^ Motluk, Alison (2001). "Big Bottom". New Scientist. 19 (7).
Jump up ^ Ehrlich, Paul R.; David S. Dobkin & Darryl Wheye. ""Bird Voices" and "Vocal Development" from Birds of Stanford essays". Retrieved 9 Sep 2008.
Jump up ^ Slabbekoorn, Hans, Smith, Thomas B. "Bird song, ecology and speciation." Philosophical Transactions: Biology Sciences 357.1420 (2002). 493-503.
Jump up ^ Zuberbühler, Klause. "Predator-specific alarm calls in Campbell's monkeys, Cercopithecus campbelli." Behavioral Ecology and Sociobiology 50.5 (2001). 414-442
Jump up ^ Boughman, Janette W. "Vocal learning by greater spear-nosed bats." Proceedings: Biological Sciences 265.1392 (1998). 227-233
Jump up ^ Jabr, Ferris (12 May 2017). "Can Prairie Dogs Talk?". The New York Times.
Jump up ^ Krulwich, Robert. "New Language Discovered: Prairiedogese". Retrieved 20 May 2015.
Jump up ^ DeMello, Margo (2007). "Yips, barks and chirps: the language of prairie dogs". Retrieved 20 May 2015.
Jump up ^ "Prairie dogs' language decoded by scientists". CBC News. 21 June 2013. Retrieved 20 May 2015.
Jump up ^ Carey, Bjorn. Whales Found to Speak in Dialects. Live Science. 3 Jan. 2006.
^ Jump up to: a b c d Bradbury, J.W., and S.L. Vehrencamp. Principles of Animal Communication. Sunderland, MA: Sinauer Associates Inc., 2011. Print.
Jump up ^ Brown, G.E.; Chivers, D.P.; Smith, R.J.F. (1995). "Localized defecation by pike: A response to labelling by cyprinid alarm pheromone?". Behavioral Ecology and Sociobiology. 36 (2): 105–110. doi:10.1007/s002650050130.
Jump up ^ Gosling, L. Morris, and S. Craig Roberts. "Scent-marking by male mammals: cheat-proof signals to competitors and mates." Advances in the Study of Behavior 30 (2001): 169-217.
Jump up ^ du P. Bothma, J.; Richet, E.A.N. le. "Evidence of the use of rubbing, scent-marking andscratching-posts by Kalahari leopards". Journal of Arid Environments. 29 (4): 511–517. Bibcode:1995JArEn..29..511D. doi:10.1016/s0140-1963(95)80023-9.
Jump up ^ "Electrocommunication". Davidson College. Archived from the original on 2012-05-29. Retrieved 2011-03-03.
Jump up ^ Bradbury, J.W., and S.L. Vehrencamp. Principles of Animal Communication. Sunderland, MA: Sinauer Associates Inc., 2011.
Jump up ^ Narins, Peter M. "Seismic Communication in Anuran Amphibians." BioScience40.4 (1990): 268. Print.
Jump up ^ (Kardong & Mackessy 1991)
Jump up ^ (Krochmal et al. 2004)
Jump up ^ (Pough et al. 1992)
^ Jump up to: a b (Gracheva et al. 2010)
Jump up ^ Kürten, L., Schmidt, U. and Schäfer, K. (1984). "Warm and cold receptors in the nose of the vampire bat, Desmodus rotundus". Naturwissenschaften. 71: 327–328. Bibcode:1984NW.....71..327K. doi:10.1007/BF00396621. CS1 maint: Multiple names: authors list (link)
Jump up ^ "Web of Life:Vibrational communication in animals". Retrieved 8 December 2012.
Jump up ^ Tauzin, Tibor; Csík, Andor; Kis, Anna; Topál, József (2015). "What or where? The meaning of referential human pointing for dogs (Canis familiaris)". Journal of Comparative Psychology. 129 (4): 334–338. doi:10.1037/a0039462. ISSN 1939-2087.
Jump up ^ Sean Senechal: Dogs can sign, too. A breakthrough method of teaching your dog to communicate to you, 2009, Random House/Crown/TenSpeed Press
Jump up ^ discussed at length by Richard Dawkins under the subject of his book The Selfish Gene
^ Jump up to: a b V. M. Janik, L. S. Sayigh, and R. S. Wells: "Signature whistle shape conveys identity information to bottlenose dolphins", Proceedings of the National Academy of Sciences, vol. 103 no 21, May 23, 2006
Jump up ^ Hare, B.; Call, J.; Tomasello, M. (1998). "Communication of food location between human and dog (Canis familiaris)". Evolution of Communication. 2: 137–159. doi:10.1075/eoc.2.1.06har.
Jump up ^ K. Guo, K. Meints, C. Hall, S. Hall & D. Mills: "Left gaze bias in humans, rhesus monkeys and rhesus domestic dogs." "Animal Cognition", vol. 12, 2009
Jump up ^ "Do animals have language? - Michele Bishop". TED Ed. 10 September 2015. Retrieved 11 September 2015.
Jump up ^ The Times May 18, 2006, p.3
Jump up ^ Brandon Kiem. "Rudiments of Language Discovered in Monkeys". Wiredscience. Retrieved 2013-03-15.
Jump up ^ Real, Leslie A. (1994-12-15). Behavioral Mechanisms in Evolutionary Ecology. University of Chicago Press. ISBN 9780226705958.
External links[edit]
Wikimedia Commons has media related to Animal communication.
Animal Communicator - Documentary
Zoosemiotics: animal communication on the web
The Animal Communication Project
International Bioacoustics Council research on animal language.
Animal Sounds different animal sounds to listen and download.
The British Library Sound Archive contains over 150,000 recordings of animal sounds and natural atmospheres from all over the world.
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Ralph Breaks the Internet
From Wikipedia, the free encyclopedia
Jump to navigation Jump to search
Ralph Breaks the Internet
Teaser poster with original title
Directed by
Rich Moore
Phil Johnston
Produced by
Clark Spencer
Screenplay by
Phil Johnston[1]
Pamela Ribon[2]
Based on
Characters
by Rich Moore
Phil Johnston
Jim Reardon
Starring
John C. Reilly
Sarah Silverman
Jack McBrayer
Jane Lynch
Taraji P. Henson
Alan Tudyk
Gal Gadot
Music by
Henry Jackman
Cinematography
Nathan Warner
Production
company
Walt Disney Pictures
Walt Disney Animation Studios
Distributed by
Walt Disney Studios
Motion Pictures
Release date
November 21, 2018 (2018-11-21)
Country
United States
Language
English
Ralph Breaks the Internet is an upcoming American 3D computer-animated comedy film produced by Walt Disney Animation Studios. The sequel to the 2012 film Wreck-It Ralph, and the 57th Disney animated feature film, it features the voices of John C. Reilly, Sarah Silverman, Jack McBrayer, Jane Lynch, Taraji P. Henson, Alan Tudyk and Gal Gadot. The film is scheduled to be released in the United States on November 21, 2018, by Walt Disney Pictures.
Contents
1 Premise
2 Voice cast
3 Popular culture cameos and references
4 Production
5 Release
5.1 Marketing
6 References
7 External links
Premise[edit]
Six years after the events of the first film,[3] the story centers on Wreck-It Ralph's adventures in the Internet data space when a Wi-Fi router gets plugged into the arcade by Mr. Litwak, as he must find a replacement for the broken steering wheel[4] of Sugar Rush. Along the way, Ralph and his best friend Vanellope von Schweetz encounter new customs, worlds, and characters, such as trendy algorithms and the Disney Princess lineup with The Muppets, Star Wars, Disney Animation, Marvel Comics and Pixar characters.[5]
Voice cast[edit]
John C. Reilly as Wreck-It Ralph, a gigantic but soft-hearted man who is the villain of the fictional arcade game Fix-It Felix Jr.[6]
Sarah Silverman as Vanellope von Schweetz, a racer/glitch in Sugar Rush who turned out to be its lost princess.[7]
Jack McBrayer as Fix-It Felix Jr., a repairman who is the hero of Fix-It Felix Jr., as well as the husband of Sergeant Calhoun.[8]
Jane Lynch as Sergeant Tamora Jean Calhoun, the lead character of Hero's Duty and Fix-It Felix Jr.'s wife.[8]
Taraji P. Henson as Yesss, an algorithm that determines the trending videos on BuzzTube.[9][10]
Alan Tudyk as KnowsMore, a search engine. Tudyk previously voiced King Candy in Wreck-It Ralph.[10][11]
Gal Gadot as Shank, a racer in Slaughter Race.[12]
James Corden[8]
Ana Ortiz[13]
Susan Lucci as Susan[14]
Bill Hader as JP Spamly.[15]
Hamish Blake as Pyro[16]
All of the Disney Princesses appear,[17][18] including: Jennifer Hale as Cinderella,[19] Kate Higgins as Aurora,[19] Jodi Benson as Ariel, Paige O'Hara as Belle, Linda Larkin as Jasmine, Irene Bedard as Pocahontas, Ming-Na Wen as Mulan, Anika Noni Rose as Tiana, Mandy Moore as Rapunzel, Kelly Macdonald as Merida, Kristen Bell as Anna, Idina Menzel as Elsa and Auliʻi Cravalho as Moana, all of whom reprise their roles from the films and other media,[19] while screenwriter Pamela Ribon will voice Snow White.[17][18][20][21][22]
Popular culture cameos and references[edit]
Similar to the first film, which included a number of cameos and references to video games, Ralph Breaks the Internet has additional features to Internet culture and to various Disney properties, including their own films, Pixar films, the Star Wars franchise, Marvel Comics, and The Muppets. Eeyore, C-3PO, R2-D2, Yoda, First Order Stormtroopers, and Iron Man will appear in the film as well.[23][24]
Production[edit]
In October 2012, director Rich Moore said that he and Disney had ideas about a sequel that would bring the characters up to date and explore online gaming and console gaming.[25] Moore stated that many of the crew and voice cast were open to the sequel, believing that they have "barely scratched the surface" of the video game world they envisioned. He also stated that he planned to include Mario and Tron in the sequel.[26][27] In 2014, the film's composer Henry Jackman said that a story for the sequel was being written.[28] In July 2015, John C. Reilly said he had signed on to reprise his role of Ralph in a projected sequel.[6]
On March 24, 2016, Moore stated that a sequel was still being planned. Moore specifically stated that a sequel would include an appearance from Mario, citing a "good relationship with Nintendo".[29] On June 30, 2016, Walt Disney Animation Studios announced that the sequel would be released on March 9, 2018, with Reilly, Moore and writer Phil Johnston attached, and that it would focus on "Ralph leaving the arcade and wrecking the Internet".[30] On December 2016, Alan Tudyk confirmed his return in the sequel.[10][31]
The sequel's title was officially announced in March 2017 as Ralph Breaks the Internet: Wreck-It Ralph 2. Further, Disney had confirmed that Reilly, Jane Lynch, Jack McBrayer, and Sarah Silverman are set to reprise their roles, while Moore returns as director joined by the first film's co-writer, Phil Johnston, in his directing debut in an animated film and Clark Spencer returns as producer.[32] In July 2018, Disney shortened the film's title to simply Ralph Breaks the Internet.[33] In August 2018, actress Gal Gadot joined the film.[34]
Release[edit]
On June 30, 2016, Walt Disney Animation Studios initially announced that the sequel, titled Ralph Breaks the Internet: Wreck-It Ralph 2, would be released on March 9, 2018.[30] However, in April 2017, A Wrinkle in Time took over its date, and the film was pushed back.[35] In July 2018, Disney shortened the film's title to simply Ralph Breaks the Internet.[33] The film will be released on November 21, 2018 in 3D, 2D and IMAX 3D.[36]
Marketing[edit]
A new poster for the film was released on February 26, 2018. Two days later, a teaser trailer for the film was released on February 28, 2018, and it quickly became viral, getting more than 4.5 million views in 24 hours.[37] A second trailer was released on June 4, 2018 with the Daft Punk song "Harder, Better, Faster, Stronger".[24] A sneak peek of the film was released on August 10, 2018.[38]
References[edit]
Jump up ^ Robinson, Laurann (February 27, 2018). "Oscars Preview: What goes in to making an Oscar-winning animated film?". KETV. Retrieved February 27, 2018.
Jump up ^ Serrao, Nivea (November 4, 2016). "Inside Slam! the new comic series all about roller derby". Entertainment Weekly. Archived from the original on November 7, 2016. Retrieved November 16, 2016.
Jump up ^ "Exclusive: 'Wreck-It Ralph 2' Director Rich Moore Teases Internet Setting, Returning Characters, and More". Collider. June 30, 2016. Archived from the original on July 3, 2016.
Jump up ^ "Disney Shares New 'Ralph Breaks the Internet: Wreck-It Ralph 2' Scenes at Annecy". Variety. June 15, 2018. Retrieved June 15, 2018.
Jump up ^ Anderton, Ethan (July 14, 2017). "'Wreck-It Ralph 2' Brings Disney Princesses Together with Marvel & Star Wars (D23 Expo 2017)". /Film. Archived from the original on July 16, 2017. Retrieved July 21, 2017.
^ Jump up to: a b Otterson, Joe (July 13, 2015). "John C. Reilly Says He Will Star in 'Wreck-It Ralph' Sequel". TheWrap. Archived from the original on July 14, 2015. Retrieved July 13, 2015.
Jump up ^ "Sequel to "Wreck-It Ralph" Hits Theaters on March 9, 2018". The Walt Disney Company. June 30, 2016. Archived from the original on August 23, 2016.
^ Jump up to: a b c Radish, Christina (October 31, 2016). "Jane Lynch on 'Mascots', Returning for 'Wreck-It Ralph 2' and More". Collider. Archived from the original on December 10, 2016. Retrieved December 6, 2016.
Jump up ^ "Taraji P. Henson Joins 'Wreck-It Ralph' Sequel, Footage Shown at D23". Variety. July 14, 2017. Retrieved August 23, 2017.
^ Jump up to: a b c Truitt, Brian (May 30, 2018). "See exclusive first photos of Taraji P. Henson, Disney princesses in 'Wreck-It Ralph 2'". USA Today. Retrieved May 30, 2018.
Jump up ^ Han, Angie (December 5, 2016). "Alan Tudyk Will Be Back for 'Frozen 2' and 'Wreck-It Ralph 2', Wants In On Marvel's 'Guardians of the Galaxy'". /Film. Archived from the original on December 6, 2016. Retrieved December 5, 2016.
Jump up ^ D'Alessandro, Anthony (August 10, 2018). "Gal Gadot Buckles Up For Disney's 'Ralph Breaks The Internet'". Deadline Hollywood. Retrieved August 10, 2018.
Jump up ^ Sneider, Jeff (February 28, 2017). "Ana Ortiz has joined the voice cast of Disney's Wreck-It Ralph 2". The Tracking Board. Archived from the original on March 1, 2017.
Jump up ^ Carter, Brooke (May 25, 2018). "Susan Lucci Net Worth 2018". Gazette Review. Retrieved May 26, 2018.
Jump up ^ Rich, Katey (June 28, 2018). "The Emmy Contenders Giving Us Hope". Little Gold Men (Podcast). Vanity Fair. Retrieved June 29, 2018.
Jump up ^ "Hamish Blake lands voice role in upcoming Disney movie 'Ralph Breaks the Internet'". Nine Network Australia. Retrieved 29 August 2018.
^ Jump up to: a b Holmes, Adam (July 14, 2017). "Wreck-It Ralph 2 Is Bringing The Original Disney Princesses Back". CinemaBlend. Archived from the original on July 16, 2017. Retrieved July 14, 2017.
^ Jump up to: a b Breznican, Anthony (July 14, 2017). "Wreck-It Ralph sequel will unite the Disney princesses — and Star Wars!". Entertainment Weekly. Archived from the original on July 15, 2017. Retrieved July 15, 2017.
^ Jump up to: a b c Johnson, Zach (June 4, 2018). "Disney Princesses Meet Vanellope von Schweetz in Wreck It Ralph 2 Trailer". E! News. Retrieved August 10, 2018.
Jump up ^ NCZ (June 4, 2018). "Wreck-It Ralph 2 Trailer". Behind The Voice Actors. Retrieved June 15, 2018.
Jump up ^ Ribon, Pamela [@PamelaRibon] (June 4, 2018). "You guys. I finally get to tell you. I'm Snow White" (Tweet). Retrieved June 4, 2018 – via Twitter.
Jump up ^ Outlaw, Kofi (July 14, 2017). "'Wreck-It Ralph 2' Bringing Back Original Disney Princess Voice Actresses". Comic Book. Archived from the original on July 16, 2017. Retrieved July 15, 2017.
Jump up ^ Carbone, Gina (July 14, 2017). "'Wreck-It Ralph 2' Brings All of the Disney Princesses Together (With C-3PO)". Moviefone. Archived from the original on July 16, 2017. Retrieved July 15, 2017.
^ Jump up to: a b Alexander, Julia (June 4, 2018). "Wreck-It Ralph 2 trailer mocks Google's annoying predictive search". Polygon. Retrieved June 4, 2018.
Jump up ^ Disney Wreck-It Ralph Director Rich Moore Is a Huge Gamer. GamerHub Videos. October 25, 2012. Event occurs at 4:22. Archived from the original on March 3, 2013. Retrieved October 25, 2012 – via YouTube.
Jump up ^ Dekel-Daks, Tal (February 8, 2013). "'Wreck-It Ralph' director Rich Moore wants Nintendo's Mario for sequel". Digital Spy. Archived from the original on September 27, 2013. Retrieved September 24, 2013.
Jump up ^ Vejvoda, Jim (February 4, 2013). "Wreck-It Ralph Director Wants Tron in the Sequel". IGN. Archived from the original on September 27, 2013. Retrieved September 24, 2013.
Jump up ^ Roberts, Sheila (April 1, 2014). "Composer Henry Jackman Talks Captain America: The Winter Soldier, His Influences, Wreck-It Ralph 2, The Interview, and More". Collider. Archived from the original on June 12, 2016. Retrieved April 3, 2014. I can't tell you more, not because I'm being coy, but I believe that it is officially on the cards. I don't know any more other than a story is indeed being written.
Jump up ^ Tilly, Chris (March 24, 2016). "Wreck-It Ralph 2 Still Happening, Might Feature Mario". IGN. Archived from the original on March 24, 2016. Retrieved March 24, 2016.
^ Jump up to: a b Snetiker, Marc (June 30, 2016). "Wreck-It Ralph 2 officially announced at Disney". Entertainment Weekly. Archived from the original on June 30, 2016. Retrieved June 30, 2016.
Jump up ^ Han, Angie (December 5, 2016). "Alan Tudyk Will Be Back for 'Frozen 2' and 'Wreck-It Ralph 2', Wants In On Marvel's 'Guardians of the Galaxy'". /Film. Archived from the original on December 6, 2016. Retrieved December 5, 2016.
Jump up ^ O'Brien, Lucy (March 28, 2017). "Wreck-It Ralph 2 is Called Ralph Breaks the Internet". IGN. Archived from the original on March 29, 2017. Retrieved March 28, 2017.
^ Jump up to: a b McClintock, Pamela (July 10, 2018). "Disney Pushes 'Indiana Jones 5' a Year to 2021, Dates 'Maleficent 2,' 'Jungle Cruise'". The Hollywood Reporter. Archived from the original on July 10, 2018. Retrieved July 10, 2018.
Jump up ^ D'Alessandro, Anthony (August 10, 2018). "Gal Gadot Buckles Up For Disney's 'Ralph Breaks The Internet'". Deadline Hollywood. Retrieved August 10, 2018.
Jump up ^ McClintock, Pamela (April 25, 2017). "'Star Wars: Episode IX' Sets Summer 2019 Release Date". The Hollywood Reporter. Archived from the original on April 26, 2017. Retrieved April 25, 2017.
Jump up ^ Lieberman, David (February 22, 2017). "Disney Films To Show On Imax Through 2019 With New Distribution Deal". Archived from the original on February 23, 2017. Retrieved March 2, 2017.
Jump up ^ Disney [@Disney] (February 27, 2018). ""There's no one I'd rather be than me." 😇😡" (Tweet). Retrieved February 27, 2018 – via Twitter.
Jump up ^ Bonomolo, Cameron (August 10, 2018). "New 'Wreck-It Ralph 2' Trailer Features Gal Gadot and More Disney Princesses". Movies. Retrieved August 12, 2018.
External links[edit]
Official website
Ralph Breaks the Internet on IMDb
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Melody Time (1948)
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Alice in Wonderland (1951)
Peter Pan (1953)
Lady and the Tramp (1955)
Sleeping Beauty (1959)
One Hundred and One Dalmatians (1961)
The Sword in the Stone (1963)
The Jungle Book (1967)
The Aristocats (1970)
Robin Hood (1973)
The Many Adventures of Winnie the Pooh (1977)
The Rescuers (1977)
The Fox and the Hound (1981)
The Black Cauldron (1985)
The Great Mouse Detective (1986)
Oliver & Company (1988)
The Little Mermaid (1989)
The Rescuers Down Under (1990)
Beauty and the Beast (1991)
Aladdin (1992)
The Lion King (1994)
Pocahontas (1995)
The Hunchback of Notre Dame (1996)
Hercules (1997)
Mulan (1998)
Tarzan (1999)
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Dinosaur (2000)
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Atlantis: The Lost Empire (2001)
Lilo & Stitch (2002)
Treasure Planet (2002)
Brother Bear (2003)
Home on the Range (2004)
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